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Russian Entomol. J. 11(3): 253263

ї RUSSIAN ENTOMOLOGICAL JOURNAL, 2002

New and little known species of Lycidae (Coleoptera) from China Новые и малоизвестные виды Lycidae (Coleoptera) из Китая Sergey V. Kazantsev С.В. Казанцев
Donetskaya Str. 13-326, Moscow 109651 Russia. ул. Донецкая 13-326 Москва 109651 Россия.

KEY WORDS: Coleoptera, Lycidae, new species, Palaearctic, China. КЛЮЧЕВЫЕ СЛОВА: Coleoptera, Lycidae, новые виды, Палеарктика, Китай. ABSTRACT.Seventeen new species of Lycidae are described from China:Macrolycus gansuensis, M. oreophilus, M. multicostatus, M. wrasei, Lycostomus tsinlingensis, L. potanini, Conderis pseudoconderoides, C. wolongensis, Cautires procautiroides, C. callimarginatus, C. miaoensis, C. leei, C. tengjiensis, Xylobanus minshanicus, X. austrogansuensis, X. macrolycoides and Libnetis taiwanus spp.n. A key to the Chinese species of the genus Macrolycus (group with parameres of the aedeagus) is provided. Additional information is given on Xylobanellus gansuensis Kazantsev, 2000. РЕЗЮМЕ.Описывается семнадцать новых видов Lycidae из Китая: Macrolycus gansuensis, M. oreophilus,M. multicostatus,M. wrasei, Lycostomus tsinlingensis, L. potanini, Conderis pseudoconderoides, C. wolongensis, Cautires procautiroides, C. callimarginatus, C. miaoensis, C. leei, C. tengjiensis, Xylobanus minshanicus, X. austrogansuensis, X. macrolycoides и Libnetis taiwanus spp.n. Дается определительная таблица китайских видов рода Macrolycus (группа с парамерами эдеагуса). Приводится дополнительная информация по Xylobanellus gansuensis Kazantsev, 2000. information on a poorly known taxon is to a large extent based on the material collected in 2001 and 2002 in remnants of primary forests of the Minshan Mountains in Southern Gansu and Northern Sichuan.

The following abbreviations are used in the paper: NME Naturhistorisches Museum, Erfurt; NTU Coleoptera Collection, National Taiwan University, Taipei; ZIN Zoological Institute, Saint-Petersburg; ZMMU Zoological Museum of Moscow University; SVK authors collection.

Material and Methods
Material used in this study was collected by hand-picking or by sweeping and beating the low strata foliage and grass, occasionally at lights. When extracted from storage bottles male specimens were dissected, with their genitalia pulled out and affixed with water-soluble glue on cardboard plates or put in microvials with glycerine.

Descriptions
Macrolycus gansuensis sp.n. Figs. 13.
DESCRIPTION. Dark brown. Pronotum, scutellum and elytra brick red. First to fourth antennomeres interiorly light brown. MALE. Head dorsally almost flat, antennal prominence rather inconspicuous, antennal cavities separated by 3 times less than their width. Eyes relatively small (separated medially above by 3.75 times their radius). Distal joint of maxillary palpi relatively small, elongate, widest in basal fourth. Antennae from 3rd joint flattened, lamellate (Fig. 1), reaching half of elytra; all joints with short decumbent pubescence. Pronotum transverse, 1.5 times wider than long, with rounded anterior angles, almost straight sides and acute prominent posterior angles, anterior margin relatively little produced forward, with conspicuous median emargination. Scutellum square, straight at apex. Elytra long (3.1 times as long as wide humerally), widest in middle, with second and fourth primary costae definitely stouter, with third rather inconspicuous, except at base. Pubescence relatively long and dense, completely hiding elytral reticulation. Legs with wide femorae and tibiae; each claw with minute tooth reaching its middle.

Introduction
The net-winged beetles of continental China have been rather intensively studied since 1996 when two first papers appeared after several decades of neglect [Bocak, 1996; Kazantsev, 1996]. Though afterwards quite a number of articles dealing with the lycid fauna of China were published, almost every new collecting trip to the area brings new species, including in the genera that have already been reviewed, such as Macrolycus Waterhouse, 1878 [Kazantsev, 2000b], Plateros Bourgeios, 1879 [Bocakovб, 1997; Kazantsev, 2001], Libnetis Waterhouse, 1878 [Bocakovб, 2000; Kazantsev, 2000a], etc. There is little doubt that further collecting in China, especially in its central provinces, will entail the discovery of yet undescribed Lycidae. This study presenting the description of several new species that belong in seven different genera and adding


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11 4 2 23 9

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5 12

8

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14

Figs. 114. Antennal joints, hind tibiae and aedeagi of Macrolycus spp.: 13 M. gansuensis sp.n., holotype , 1 3rd and 4th antennal joints; 23 aedeagus ventrally (2) and laterally (3); 47 M. oreophilus sp.n., holotype , 4 3rd and 4th antennal joints; 5 hind tibia; 67 aedeagus ventrally (6) and laterally (7); 8 M. shaanxiensis Kazantsev, holotype , hind tibia; 911 M. multicostatus sp.n., holotype , 9 3rd and 4th antennal joints; 1011 aedeagus ventrally (10) and laterally (11); 1214 M. wrasei sp.n., holotype , 12 3rd and 4th antennal joints; 1314 aedeagus ventrally (13) and laterally (14). Scale: 0.5 mm. Figs. 114. Антенные членики, задние голени и эдеагусы Macrolycus spp.: 13 M. gansuensis sp.n., голотип, , 1 34-й антенные членики; 23 эдеагус вентрально (2) и латерально (3); 47 M. oreophilus sp.n., голотип, , 4 34-й антенные членики; 5 задняя голень; 67 эдеагус вентрально (6) и латерально (7); 8 M. shaanxiensis Kazantsev, голотип, , задняя голень; 911 M. multicostatus sp.n., голотип, , 9 34-й антенные членики; 1011 эдеагус вентрально (10) и латерально (11); 1214 M. wrasei sp.n., голотип, , 12 34-й антенные членики; 1314 эдеагус вентрально (13) и латерально (14). Масштаб: 0,5 мм.

Holotype : China: S Gansu, Tepo (Tewo), 25002800 m, 2628.VI.2001, S. Kazantsev (SVK); paratype : same label; paratype : China: S Gansu, Tsagan (70 km W Wudu), 2300 2400 m, 1617.VI.2001, S. Kazantsev (SVK).

Aedeagus with relatively broad median piece and short and wide parameres (Figs. 23). Length: 13.5 mm. Width (humerally): 3.5 mm. FEMALE. Unknown.

DIAGNOSIS. M. gansuensis sp.n. belongs in the group with noticeable parameres of the aedeagus (see Kazantsev, 2000b), at the same time readily distinguishable from its other members by the almost straight anterior pronotal margin notched medially, shorter antennal lamellae (Fig. 1) and more robust and flattened basally median piece of the aedeagus with shorter and wider parameres (Figs. 23).


New and little known species of Lycidae (Coleoptera) from China Macrolycus oreophilus sp.n. Figs. 47.
DESCRIPTION. Black. Pronotum and elytra bright red. MALE. Head dorsally almost flat, antennal prominence rather inconspicuous, antennal cavities separated by 5 times less than their width. Eyes relatively small (separated medially above by 3.2 times their radius). Distal joint of maxillary palpi large, elongate, widest in basel. Antennae from 3rd joint flattened, lamellate (Fig. 4), reaching over half of elytra; all joints with short decumbent pubescence. Pronotum transverse, about 1.5 times wider than long, with pronounced blunt anterior angles, almost straight sides and long acute posterior angles, anterior margin triangularly produced forward. Scutellum elongate, parallel-sided, straight at apex. Elytra long (4.3 times longer than wide humerally), widest just behind middle, with second and fourth primary costae definitely stouter, with third rather inconspicuous. Pubescence relatively long and dense, almost hiding irregular elytral punctuation. Legs with straight narrow posterior tibiae (Fig. 5). Aedeagus with slender median piece and parameres, and narrow dorso-apical opening of the median piece (Figs. 67). Length: 12.415.5 mm. Width (humerally): 2.43.1 mm. FEMALE. Similar to male, but broader, with just strongly serrate antennae.

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Macrolycus multicostatus sp.n. Figs. 911.
DESCRIPTION. Dark brown. Pronotum and elytra dark red. MALE. Head dorsally almost flat, antennal prominence rather inconspicuous, antennal cavities separated by 3 times less than their width. Eyes small (separated medially above by 4 times their radius). Distal joint of maxillary palpi large, considerably exceeding in length and width all other joints taken together, twice as long as wide, widest at base. Antennae from 3rd joint flattened, from 4th lamellate (Fig. 9), reaching over half of elytra; all joints with short decumbent pubescence. Pronotum almost square, only 1.1 times wider than long, with pronounced blunt anterior angles, widening anteriorly sides and small acute posterior angles, anterior margin triangularly produced forward. Scutellum square, straight at apex. Elytra long (3.6 times as long as wide humerally), widest just behind middle, with second and fourth primary costae definitely stouter; interstices with double rows of reticulation, arranged in basal half in irregular cells. Pubescence relatively short and scarce, not consealing elytral reticulation. Legs with narrow femorae and tibiae; each claw with tooth almost reaching its apex. Aedeagus with slender straight median piece and relatively wide roundish dorso-apical opening and long lateral rib (Figs. 1011). FEMALE. Similar to male, but antennae without lamellae, pronotum slightly wider than elytra basally, femorae and tibiae definitely wider and tarsal claws with shorter teeth. Length: 11.112.0 mm. Width (humerally): 2.52.9 mm.
Holotype : China: W Sichuan, Jiulong Co., Taka He valley, 25004000 m, 28њ4529њ09(N) & 101њ4250(E), 210.VII.2001, L. & R. Businsky (SVK); paratype : same label (SVK).

DIAGNOSIS. M. oreophilus sp.n. also belongs in the same group as M. gansuensis sp.n., is similar in certain aspects to M. shaanxiensis Kazantsev, differing from it by the narrow and straight hind tibiae (Fig. 5) and the details of the aedeagus (Figs. 67). VARIATION. Paratype specimens from western Sichian differ in having dark red coloration of the upperside.

Holotype : CHINA: N YUNNAN, Haba Mts, Bailakou Pass, 3450 m, VII.10.2002, S.Murzin leg. (SVK); paratypes, 4 : same label; paratype, : China: YUNNAN, Yulongxue Shan, 3600 m, 1416.VI.1998, S.Murzin leg.; paratypes, and 2 : China: W Sichuan, Jiulong Co., Taka He valley, 2500 4000 m, 28њ4529њ09(N) & 101њ4250(E), 210.VII.2001 coll. L. & R. Businsky (SVK).

The difference between the 4 currently known Chinese Macrolycus species belonging in the group with the parameres of the aedeagus is shown in the key below: A
KEY TO

DIAGNOSIS. M. multicostatus sp.n. somewhat resembles M. aurantiacus Kazantsev in having a non-lamellate third antennal segment (Fig. 9), but can be easily differentiated from almost all congenerics, including M. aurantiacus, by the presence of secondary elytral costae dividing each interstice into two rows of irregular cells and the details of the aedeagus (Figs. 1011). M. yunnanus Kazantsev, the only Macrolycus species that has similar elytral structure, belongs in a different group characterized by the presence of parameres [Kazantsev, 2000b].

Macrolycus wrasei sp.n. Figs. 1214.
DESCRIPTION. Dark brown. Anterior and lateral margins of pronotum and elytra dark red, with purplr-red pubescence. MALE. Head dorsally almost flat, antennal prominence rather inconspicuous, antennal cavities separated by 5 times less than their width. Eyes relatively small (separated medially above by 3.7 times their radius). Distal joint of maxillary palpi large, about twice as long as wide, widest at base, considerably exceeding in length and width all other joints taken together. Antennae from 3rd joint flattened and lamellate (Fig. 12), reaching two thirds of elytra; all joints with short decumbent pubescence. Pronotum transverse, 1.25 times wider than long, widening anteriorly, with pronounced acute anterior and prominent acute (mostly propleurical) posterior angles, anterior margin bisinuate. Scutellum square, slightly concave at apex. Elytra long (about 3.9 times longer than wide humerally), widest in distal third, with third primary costa definitely less pronounced in distal half; interstices with dense fine punctu-

C

HINESE SPECIES OF THE GENUS

PARAMERES OF THE AEDEAGUS

)

M

ACROLYCUS (WITH

1. Upperside uniformly orange. Elytra with irregular double rows of cells in interstices ....... M. yunnanus Kazantsev Elytra red. Elytral interstices with dense irregular punctuation ............................................................................... 2 2. Third antennal joint triangular. Posterior pronotal angles inconspicuous. Aedeagus Figs. 23 .......................... ......................................................... M. gansuensis sp.n. Third antennal joint lamellate. Posterior pronotal angles conspicuously produced laterlally ................................ 3 3. Posterior tibiae relatively broad and curved (Fig. 8) ....... ............................................. M. shaanxiensis Kazantsev Posterior tibiae narrow and straight (Fig. 5). Median lobe of aedeagus with narrow dorsal opening (Fig. 6) .......... ......................................................... M. oreophilus sp.n.


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19 16 18

20

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24 22 23 25 26

Figs. 1526. Antennal joints, rostrums and aedeagi of Lycostomus and Conderis spp.: 1517 L. tsinlingensis sp.n., holotype , 15 rostrum; 1617 aedeagus ventrally (16) and laterally (17); 1820 L. potanini sp.n., holotype , 18 rostrum; 1920 aedeagus ventrally (19) and laterally (20); 2123 C. pseudoconderoides sp.n., holotype , 21 3rd and 4th antennal joints; 2223 aedeagus ventrally (22) and laterally (23); 2426 M. multicostatus sp.n., holotype , 24 3rd and 4th antennal joints; 2526 aedeagus ventrally (25) and laterally (26). Scale: 0.5 mm. Рис. 1526. Антенные членики, рострумы и эдеагусы Lycostomus и Conderis spp.: 1517 L. tsinlingensis sp.n., голотип, , 15 рострум; 1617 эдеагус вентрально (16) и латерально (17); 1820 L. potanini sp.n., голотип, , 18 рострум; 19 20 эдеагус вентрально (19) и латерально (20); 2123 C. pseudoconderoides sp.n., голотип, , 21 34-й антенные членики; 2223 эдеагус вентрально (22) и латерально (23); 2426 C. wolongensis sp.n., голотип, , 24 34-й антенные членики; 2526 эдеагус вентрально (25) и латерально (26). Масштаб: 0,5 мм.

Holotype : China (N-Sichuan), Daba Shan, 65 km N Fengjie, river bank, 31њ40N/109њ34E, ~1000 m, (fine gravel bank), 10.VII.2001, Wrase (08) (SVK).

ation. Pubescence short and dense, almost consealing elytral structure. Legs with relatively narrow tibiae. Aedeagus with roundish dorso-apical cavity open distally and slightly dorso-ventrally dilated apex of median lobe (Figs. 1314). FEMALE. Unknown. Length: 13.9 mm. Width (humerally): 3.0 mm.

Lycostomus tsinlingensis sp.n. Figs. 1517.
DESCRIPTION. Black. Pronotal and elytral margins cinnobar red. MALE. Head dorsally with conspicuous double impression behind antennal prominence, antennal cavities separated by minute lamina. Eyes relatively small (separated medially above by 2.3 times their radius). Rostrum moderately long (Fig. 15). Distal joint of maxillary palpi small, elongate, 1.8 times longer than wide. Antennae from 3rd joint flattened and serrate, reaching about half of elytra; all joints with short decumbent pubescence. Pronotum semioval, transverse, 1.6 times wider than long, with rounded anterior and prominent acute posterior angles. Scutellum trapezoidal, almost straight at apex.

DIAGNOSIS. M. wrasei sp.n. may be placed near M. murzini Kazantsev from South Shaanxi (Taibaishan) due to the similar structure of the pronotum and the aedeagus, but is readily distinguished by the coloration, the differently shaped apex of the median lobe and other details of the aedeagus (Figs. 1314).


New and little known species of Lycidae (Coleoptera) from China
Elytra long (3.3 times as long as wide humerally), widened posteriorly, widest in distal fourth, with second and fourth primary costae slightly stouter in distal half; interstices with distinct fine irregular reticulation. Pubescence relatively short and scarce, not consealing elytral structure. Legs with narrow femorae and tibiae; tibiae conspicuously curved. Aedeagus with relatively long parameres and short median piece (Figs. 1617). FEMALE. Similar to male, but antennae shorter and less serrate. Length: 11.016.5 mm. Width (humerally): 2.94.6 mm.

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Conderis pseudoconderoides sp.n. Figs. 2123.
DESCRIPTION. Black. Elytra except at base, suture and margins dark red. Pronotal margins and ribs, as well as elytral base, suture and margins with reddish pubescence. MALE. Head dorsally almost flat, antennal prominence rather inconspicuous, antennal cavities separated by 4 times less than their width. Eyes relatively small (separated medially above by 3 times their radius). Distal joint of maxillary palpi about as long as 2nd, slightly longer than wide, widest near base. Antennae from 3rd joint flattened and serrate (Fig. 21), reaching over half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum transverse, 1.3 times wider than long, trapezoidal, with pronounced slightly rounded anterior and acute produced anterior-laterally posterior angles; rounded anteriorly and conspicuously bisinuate posteriorly; median cell posteriorly closed, connected with posterior margin by half cell long rib. Scutellum trapezoidal, almost straight at apex. Elytra long (4.1 times as long as wide humerally), slightly widening posteriorly, with almost equally developed primary costae, only 4th noticeably stouter in all its length except at base; interstices with double rows of distinct and rather irregular cells. Pubescence represented by minute hairs distributed along elytral costae. Legs slender and narrow. Aedeagus Figs. 2223. FEMALE. Similar to male, but antennae serrate, without lamellae. Length: 7.011.1 mm. Width (humerally): 1.62.8 mm.
Holotype : China: S Gansu, Tsagan (70 km W Wudu), 23002400 m, 1617.VI.2001, S. Kazantsev (SVK); paratypes, 2 and 1 : same label; paratype : China: Shaanxi, Taibaishan Mts, env. Haozhenzi, 15002000 m, 33њ52N 107њ44E, VI VII.2000, A. Plutenko leg. (SVK).

Holotype : China: Shaanxi, env. Haozhenzi, 13502000 m, 1424.VI.(19)99, S. Murzin leg. (SVK); paratypes, 2 and 2 : same label; paratype : China: Shaanxi, Taibaishan Nat Park, 1350 m, 10.VI.(19)99, S. Murzin leg.; paratypes, and 3 : China: S Gansu, Tsagan (70 km W Wudu), 23002400 m, 16 17.VI.2001, S. Kazantsev; paratype : China: S Gansu, Minshan, 70 km W Wudu, 2000-2700 m, 25.VII.2000, A. Gorodinsky (SVK); paratypes, 4 and 15 : China: Shaanxi, Taibaishan Mts, env. Haozhenzi, 15002000 m, 33њ52N 107њ44E, VI VII.2000, A. Plutenko leg. (ZMMU and SVK).

DIAGNOSIS. L. tsinlingensis sp.n. is evidently closely allied to L. porphyrophorus (Solsky), both species occurring in Taibaishan (Qinling Mts, Shaanxi), differing by the coloration, longer rostrum (Fig. 15), shorter and scarcer elytral pubescence, relatively longer parameres and shorter median piece of the aedeagus (Figs. 1617). REMARKS. The dark elytral stripe may considerably vary both in size and tone.

Lycostomus potanini sp.n. Figs. 1820.
DESCRIPTION. Black. Pronotum anteriorly, in anterior half of lateral margins and in posterior angles dark red; elyrta dark brown with red lateral emargination. MALE. Head dorsally with conspicuous impression behind antennal prominence, antennal cavities separated by minute lamina. Eyes relatively small (separated medially above by 2.5 times their radius). Rostrum short (Fig. 18). Distal joint of maxillary palpi small, elongate, 1.4 times longer than wide, widest in middle. Antennae from 3rd joint flattened, from 4th serrate, reaching about half of elytra; all joints with short decumbent pubescence. Pronotum semi-oval, transverse, 1.5 times wider than long, with rounded anterior and prominent acute posterior angles. Scutellum small, trapezoidal, almost straight at apex. Elytra long (2.8 times as long as wide humerally), conspicuously widened posteriorly, widest in distal third, with third primary costa definitely weaker than others; interstices with distinct fine irregular reticulation. Pubescence relatively short and scarce, not consealing elytral structure. Legs with narrow femorae and tibiae; tibiae conspicuously curved. Aedeagus with relatively short median piece (Figs. 1920). FEMALE. Similar to male, but antennae shorter, elytra less broadened. Length: 11.912.7 mm. Width (humerally): 3.23.6 mm.

DIAGNOSIS. C. pseudoconderoides sp.n. perhaps should be put near C. longipennis (Pic), formerly placed in the genus Pseudoconderis Pic, 1921, as it has similar pronotal structure, distinct elytral reticulation and minute hairs along elytral costae, but can be easily distinguished by the coloration, serrate antennae (Fig. 21) and the details of the aedeagus (Figs. 2223).

Conderis wolongensis sp.n. Figs. 2426.
DESCRIPTION. Black. Pronotum deep brown red; elytra dark red. MALE. Head dorsally almost flat, antennal prominence rather inconspicuous, antennal cavities separated by 4 times less than their width. Eyes relatively small (separated medially above by 3.3 times their radius). Distal joint of maxillary palpi about as long as 2nd, slightly longer than wide, widest near middle. Antennae from 3rd joint flattened and inconspicuously serrate (Fig. 24), reaching over half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum transverse, 1.3 times wider than long, trapezoidal, with slightly rounded anterior and acute produced anterior-laterally posterior angles; rounded anteriorly and conspicuously bisinuate posteriorly; median cell posteriorly closing at pronotal margin. Scutellum trapezoidal, almost straight at apex. Elytra long (3.6 times as long as wide humerally), slightly widening posteriorly, with 4th noticeably stouter in all its length except at base; interstices with obscure double rows of

DIAGNOSIS. L. potanini sp.n. belongs in the same group as L. porphyrophorus and L. tsinlingensis sp.n., differing from both by the coloration and the short median piece of the aedeagus (Figs. 1920).

Holotype : Sze-tchuan, 1885, G. Potanin, Lycostomus modestus Ksw. (Bourgeois manuscript label), 244 (ZIN); paratypes, 3 and 1 : China: Sichuan, 70 km E Chengkou, 18002000 m, 17.VII.1995, A. Shamaev leg. (SVK).


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33 29 27

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31
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34

35
holotype , 27 holotype , 30 holotype , 33 голотип, , 27 голотип, , 30 голотип, , 33

Figs. 2735. Antennal joints and aedeagi of Xylobanellus and Cautires spp.: 2729 X. gansuensis Kazantsev, to 4th antennal joints; 2829 aedeagus ventrally (28) and laterally (29); 3032 C. procautiroides sp.n., nd to 4th antennal joints; 3132 aedeagus ventrally (31) and laterally (32); 3335 C. callimarginatus sp.n., nd to 4th antennal joints; 3435 aedeagus ventrally (34) and laterally (35). Scale: 0.5 mm. Рис. 2735. Антенные членики и эдеагусы Xylobanellus и Cautires spp.: 2729 X. gansuensis Kazantsev, 24-й антенные членики; 2829 эдеагус вентрально (28) и латерально (29); 3032 C. procautiroides sp.n., 24-й антенные членики; 3132 эдеагус вентрально (31) и латерально (32); 3335 C. callimarginatus sp.n., 24-й антенные членики; 3435 эдеагус вентрально (34) и латерально (35). Масштаб: 0,5 мм.

Holotype : China: Sichuan, 35 km W Wolong, Dansheng, 2800 m, 717.VII.2000, S. Murzin leg. (SVK); paratypes, and : same label (SVK).

cells. Primary costae with short dense pubescence; interstices with scarcer hairs. Aedeagus Figs. 2526. FEMALE. Similar to male, but antennae slightly less serrate and pronotum wider and relatively larger. Length: 7.79.9 mm. Width (humerally): 1.72.2 mm.

ation, pronotal structure, less conspicuously serrate antennae (Fig. 24), longer and less strictly confined to the costae elytral pubescence and the details of the aedeagus (Figs. 2526).

Xylobanellus gansuensis Kazantsev, 2000 Figs. 2729.
MALE. Differs from female in having slenderer body and more serrate antennae, with acute apices of their teeth (Fig. 27).

DIAGNOSIS. C. wolongensis sp.n. is similar in many aspects to C. pseudoconderoides sp.n., differing by the color-


New and little known species of Lycidae (Coleoptera) from China
Aedeagus Fig. 2829.

259

STUDIED MATERIAL: , China: S Gansu, Tsagan (70 km W Wudu), 23002400 m, 1617.VI.2001, S. Kazantsev (SVK).

REMARKS. As the description of this species was based on a female it is complemented with some morphological details of the opposite sex. In addition to characters mentioned in the diagnosis [Kazantsev, 2000a] the male differs from X. erythropterus (Baudi), distributed in eastern Europe through Northern Asia to the Pacific Coast, by the definitely more serrate antennae (Fig. 27), absence of a broad membranous lobe in the distal portin of the parameres and other details of the aedeagus (Fig. 2829).

Cautires procautiroides sp.n. Figs. 3032.
DESCRIPTION. Black. Elytra and lateral pronotal margins dark brown with reddish pubescence. MALE. Head dorsally with shallow impression and fine median rib behind antennal prominence, antennal cavities separated by minute lamina. Eyes large (separated medially above by 1.8 times their radius). Distal joint of maxillary palpi small, about as long as 2nd, slightly longer than wide, widest at middle. Antennae from 3rd joint lamellate (Fig. 30), reaching half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum small (9 times shorter than elytra), transverse, 1.4 times wider than long, trapezoidal, with blunt anterior and small acute posterior angles; slightly produced forward anteriorly and conspicuously bisinuate posteriorly; median cell open posteriorly; anterior and lateral ribs well developed. Scutellum parallel-sided, deeply and broadly cleft at apex. Elytra long (3.75 times as long as wide humerally), slightly widening posteriorly, with 4th primary costa noticeably stouter than others in distal half; interstices with double rows of rather irregular elongate cells, becoming one row in middle of elytra, especially noticeable in first and last interstice. Pubescence uniform, sparse and short. Legs slender and narrow. Aedeagus Figs. 3132. FEMALE. Similar to male, but antennae with shorter lamellae. Length: 9.711.0 mm. Width (humerally): 2.22.5 mm.

about 5 times their width. Eyes relatively large (separated medially above by 1.8 times their radius). Distal joint of maxillary palpi small, slightly longer than wide. Antennae from 3rd joint lamellate (Fig. 33), reaching about half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum transverse, 1.2 times wider than long, trapezoidal, with rounded anterior and small acute posterior angles; conspicuously produced forward medially at anterior margin and bisinuate posteriorly; median cell narrowly open posteriorly; anterior and lateral ribs well developed. Scutellum almost parallel-sided, deeply and broadly cleft at apex. Elytra long (3.8 times as long as wide humerally), slightly widening posteriorly, with 2nd primary costa noticeably stouter than others in distal half; interstices with regular double rows of square cells. Pubescence uniform, sparse and short. Aedeagus Figs. 3435. FEMALE. Unknown. Length: 10.0 mm. Width (humerally): 2.2 mm.
Holotype : China: W Hubei, Shennongjia Nat. Res., 2000 2200 m, 38.VI.(19)95, S. Kurbatov (SVK).

DIAGNOSIS. C. callimarginatus sp.n. can be easily distinguished from other Cautires species from China by the coloration, as well as by the details of the aedeagus (Figs. 3435).

Cautires miaoensis sp.n. Figs. 3638.
DESCRIPTION. Black. Elytra brick red. MALE. Head dorsally almost flat, shining, antennal cavities separated by minute lamina. Eyes relatively small (separated medially above by 2.75 times their radius). Distal joint of maxillary palpi small, slightly longer than wide. Antennae from 3rd joint lamellate (Fig. 36), reaching over half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum slightly transverse, 1.2 times wider than long, trapezoidal, with rounded anterior and small acute posterior angles; slightly produced forward anteriorly and bisinuate posteriorly; median cell closed at posterior margin; anterior ribs except median one feebly developed; lateral carinae hardly noticeable. Scutellum trapezoidal, slightly emarginate at apex. Elytra long (4 times as long as wide humerally), almost parallel-sided, with 2nd primary costa noticeably stouter than others in distal half; interstices with double rows of rather irregular elongate cells, nearly transferring into one row in middle of elytra, especially manifest in first and second interstices. Pubescence in basal half relatively dense distributed along primary costae, with bottom of cells hairless; in distal half pubescence almost uniform. Aedeagus Figs. 3738. FEMALE. Unknown. Length: 7.68.5 mm. Width (humerally): 1.62.0 mm.

DIAGNOSIS. C. procautiroides sp.n. can be easily distinguished from similarly coloured congenerics by the elytral retuculation with one row of cells in the middle of the elytra, especially noticeable in the first and last interstices (resembling the elytral structure characteristic of the genus Procautires Kleine, 1925) and the details of the aedeagus (Figs. 3132).

Holotype : China: S Gansu, Tsagan (70 km W Wudu), 23002400 m, 1617.VI.2001, S. Kazantsev (SVK); paratypes, 3 : same label; paratype female: China (W-Hubei), Daba Shan, pass E Mt. Da Shennongjia, 12 km NW Muyuping, 31њ30N/ 110њ21E, 1950 m, (dry creek vall./mix. decid. for.), 1622.VII.2001, Wrase (13) (NME); paratypes, 2 : China: Shaanxi, Taibaishan Mts, env. Haozhenzi, 15002000 m, 33њ52N 107њ44E, VI VII.2000, A. Plutenko leg.; paratype : China: N Sichuan, valley 5 km N Wenchuan, 1800 m, 28.VII.2001, S. Murzin leg. (SVK).

Holotype : China: Guizhou, env. Kaili-Laishan, Miao Mts, 1200 m, 1619.VI.2000, S. Murzin (SVK); paratypes, 4 : same label (SVK).

DIAGNOSIS. C. miaoensis sp.n. is distinguishable from other Chinese Cautires by the coloration, peculiar elytral reticulation and pubescence, and the details of the aedeagus (Figs. 3738).

Cautires callimarginatus sp.n. Figs. 3335.
DESCRIPTION. Black. Pronotum except at disk and external elytral emargination red. MALE. Head dorsally with inconspicuous impression behind antennal prominence, antennal cavities separated by

Cautires leei sp.n. Figs. 3941.
DESCRIPTION. Black. Elytra orange red; pronotum laterally in middle reddish-brown. MALE. Head dorsally almost flat, shining, antennal cavities separated by about 5 times their width. Eyes small


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36

39 37 38

40

41

42

43

44

45

46

47

Figs. 3647. Antennal joints and aedeagi of Cautires and Xylobanus spp.: 3638 C. miaoensis sp.n., holotype , 36 3rd and 4th antennal joints; 3738 aedeagus ventrally (37) and laterally (38); 3941 C. leei sp.n., holotype , 39 3rd to 4th antennal joints; 4041 aedeagus ventrally (40) and laterally (41); 4244 C. tengjiensis sp.n., holotype , 42 2nd to 4th antennal joints; 4344 aedeagus ventrally (43) and laterally (44); 4547 X. minshanicus sp.n., holotype , 45 3rd to 4th antennal joints; 46 47 aedeagus ventrally (46) and laterally (47). Scale: 0.5 mm. Рис. 3647. Антенные членики и эдеагусы Cautires и Xylobanus spp.: 3638 C. miaoensis sp.n., голотип, , 36 34-й антенные членики; 3738 эдеагус вентрально (37) и латерально (38); 3941 C. leei sp.n., голотип, , 39 34-й антенные членики; 4041 эдеагус вентрально (40) и латерально (41); 4244 C. tengjiensis sp.n., голотип, , 42 34-й антенные членики; 4344 эдеагус вентрально (43) и латерально (44); 4547 X. minshanicus sp.n., голотип, , 45 34-й антенные членики; 4647 эдеагус вентрально (46) и латерально (47). Масштаб: 0,5 мм.

(separated medially above by 2.6 times their radius). Distal joint of maxillary palpi small, elongate, hardly wider than preceeding joints. Antennae from 3rd joint lamellate (Fig. 39), reaching half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum 1.3 times wider than long, trapezoidal, with parallel in basal fifth and conspicuously narrowing anteriorly sides, prominent anterior and minute acute posterior angles; convex anteriorly and bisinuate posteriorly; median cell narrowly open at posterior margin; all pronotal carinae fine, but

well noticeable. Scutellum parallel-sided, broadly and deeply emarginate at apex. Elytra long (3.5 times as long as wide humerally), slightly widening posteriorly, with 2nd primary costa noticeably stouter than others in distal half; interstices with double rows of rather irregular elongate cells. Pubescence uniform, relatively dense and short. Aedeagus Figs. 4041. FEMALE. Unknown. Length: 8.9 mm. Width (humerally): 2.1 mm.


New and little known species of Lycidae (Coleoptera) from China
Holotype : Taiwan: Kaohsiung, Tengji, 22.VI.2000, C.-F. Lee leg. (NTU).

261

DIAGNOSIS. C. leei sp.n. is similar in coloration to C. miaoensis sp.n., readily distinguishable by the shape of the pronotum, elytral reticulation and details of the aedeagus (Figs. 4041).

Cautires tengjiensis sp.n. Figs. 4244.
DESCRIPTION. Black. Pronotum except at disk posteriorly and elytra reddish testaceus. MALE. Head dorsally slightly concave behind antennal prominence, antennal cavities separated by about 3 times their width. Eyes small (separated medially above by 2.6 times their radius). Distal joint of maxillary palpi small, elongate, hardly wider than preceeding joints. Antennae from 3rd joint lamellate (Fig. 42), reaching about half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum 1.3 times wider than long, with slightly convex narrowing anteriorly sides, rounded anterior and acute posterior angles; convex anteriorly and bisinuate posteriorly; median cell narrowly open at posterior margin; all pronotal carinae except mdean one hardly noticeable. Scutellum parallel-sided, rather broadly and deeply emarginate at apex. Elytra long (3.6 times as long as wide humerally), slightly widening posteriorly, with 2nd primary costa slightly stouter in distal half; interstices with double rows of regular elongate cells. Pubescence sparse and short, distributed along costae, leaving bottom of cells hairless. Aedeagus Figs. 4344. FEMALE. Unknown. Length: 7.5 mm. Width (humerally): 1.8 mm. DIAGNOSIS. C. tengjiensis sp.n. is similar in coloration to C. katoi Nakane, but in addition to the aedeagus (Figs. 43 44) can be distinguished by the shape of the pronotum with feeble lateral carinae.

DIAGNOSIS. X. tiphionus Kazantsev, smaller eyes, serrate pronotum, details of

Holotype : China: N Sichuan, Nanping, 15001800 m, 4 6.VII.2001, S. Kazantsev (SVK); paratypes, and : same label; paratypes, 2 : China: N Sichuan, valley 5 km N Wenchuan, 2000 m, 35.VII.2001, S. Murzin; paratype, : China: N Sichuan, valley 5 km N Wenchuan, 1800 m, VI.8.2002, S. Murzin (SVK).

FEMALE. Similar to male, but antennae slightly less serrate. Length: 6.57.8 mm. Width (humerally): 1.61.8 mm.

minshanicus sp.n. differs from X. mondescribed from Central Sichuan, by the male antennae (Fig. 45), shape of the the aedeagus (Figs. 4647), etc.

Xylobanus austrogansuensis sp.n. Figs. 4850.
DESCRIPTION. Black. Pronotal and elytral margins, costae and ribs red brown with reddish pubescence. MALE. Head dorsally almost flat, shining, antennal cavities separated by minute lamina. Eyes large (separated medially above by 1.2 times their radius). Distal joint of maxillary palpi small, narrow, 1.7 times longer than wide. Antennae from 3rd joint lamellate (Fig. 48), reaching basal third of elytra; all antennomeres with short dense decumbent pubescence. Pronotum nearly as wide as long, slightly narrowing anteriorly, with slightly concave sides, blunt anterior and prominent acute posterior angles; triangularly produced forward anteriorly and medially convex posteriorly; median cell closed at posterior margin; all ribs well developed. Scutellum parallel-sided, with broad deep emargination at apex. Elytra long (4.3 times as long as wide humerally), parallel-sided, with all primary costa almost equally developed; interstices with elongate cells. Pubescence scarce and short, spread along primary costae, with bottom of cells hairless. Aedeagus Figs. 4950. FEMALE. Unknown. Length: 9.5 mm. Width (humerally): 1.9 mm. DIAGNOSIS. X. austrogansuensis sp.n. differs from other Xylobanus known from China by the short lamellate antennae, elongate cells of the elytral reticulation, details of the aedeagus (Figs. 4950), etc.
Holotype : China: S Gansu, Tsagan (70 km W Wudu), 23002400 m, 1617.VI.2001, S. Kazantsev (SVK).

Holotype : Taiwan: Kaohsiung, Tengji, 22.VI.2000, C.-F. Lee leg. (NTU).

Xylobanus minshanicus sp.n. Figs. 4547.
DESCRIPTION. Black. Pronotum dark red brown with reddish pubescence at lateral margins; elytra with dark red costae and transverse ribs. MALE. Head dorsally almost flat, shining, antennal cavities separated by about 7 times their width. Eyes small (separated medially above by 2.6 times their radius). Distal joint of maxillary palpi small, stout, only slightly longer than wide. Antennae from 3rd joint serrate (Fig. 45), reaching over half of elytra; all antennomeres with short dense decumbent pubescence. Pronotum nearly as wide as long, narrowing anteriorly, with blunt anterior and prominent acute posterior angles; triangularly produced forward anteriorly and bisinuate posteriorly; median cell open at posterior margin; anterior ribs well developed; lateral carinae hardly noticeable. Scutellum parallel-sided, with broad deep emargination at apex. Elytra long (3.6 times as long as wide humerally), parallel-sided, with 2nd primary costa slightly stouter than others in distal half; interstices with regular transverse cells. Pubescence relatively dense and short, spread along primary costae, with bottom of cells hairless. Aedeagus Figs. 4647.

Xylobanus macrolycoides sp.n. Figs. 5153.
DESCRIPTION. Black. Pronotum , scutellum and elytra orange red. MALE. Head dorsally with conspicuous transverse impression behind antennal prominence, antennal cavities separated by about 6.4 times their width. Eyes relatively large (separated medially above by 1.6 times their radius). Distal joint of maxillary palpi small, slightly longer than wide. Antennae from 3rd joint serrate (Fig. 51), reaching over half of elytra; all antennomeres with short dense decumbent pubescence; joints 3 to 11 additionally with sparse minute scales. Pronotum transverse, 1.6 times wider than long, trapezoidal, with concave narrowing anteriorly sides, rounded anterior and prominent acute posterior angles; evenly produced forward anteriorly and bisinuate posteriorly; median cell open at posterior margin; anterior ribs except median one feebly developed; lateral carinae hardly noticeable. Scutellum parallel-sided, with rather broad roundish emargination at apex.


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48

49

50

51

52

53

54

55

56

Figs. 4856. Antennal joints and aedeagi of Xylobanus and Libnetis spp.: 4850 X. austrogansuensis sp.n., holotype , 48 3rd and 4th antennal joints; 4950 aedeagus ventrally (49) and laterally (50); 5153 X. macrolycoides sp.n., holotype , 51 3rd to 4th antennal joints; 5253 aedeagus ventrally (52) and laterally (53); 5456 L. taiwanus sp.n., holotype , 5456 aedeagus ventrally (54), laterally (55) and dorsally (56). Scale: 0.5 mm. Рис. 4856. Антенные членики и эдеагусы Xylobanus и Libnetis spp.: 4850 X. austrogansuensis sp.n., голотип, , 48 3 4-й антенные членики; 4950 эдеагус вентрально (49) и латерально (50); 5153 X. macrolycoides sp.n., голотип, , 51 34-й антенные членики; 5253 эдеагус вентрально (52) и латерально (53); 5456 L. taiwanus sp.n., голотип, , 5456 эдеагус вентрально (54) латерально (55) и дорсально (56). Масштаб: 0,5 мм.

Holotype : Taiwan: Kaohsiung, Tengji, 22.VI.2000, C.-F. Lee leg. (NTU); paratypes, 2 and 3 : same label; paratype, : Taiwan: Taipei, Wulai, 24.XII.1996, S.D. Lee leg. (NTU and SVK).

Elytra long (4 times as long as wide humerally), almost parallel-sided, with 2nd primary costa noticeably stouter in distal half; interstices with regular transverse cells. Pubescence relatively dense and short, distributed along primary costae, with bottom of cells hairless. Aedeagus with twisted apical portion and sclerotized process of internal sack (Figs. 5253 ). FEMALE. Similar to male, but eyes smaller and antennae less serrate. Length: 8.010.7 mm. Width (humerally): 1.92.4 mm.

Libnetis taiwanus sp.n. Figs. 5456.
DESCRIPTION. Black. Second antennal joint apically testaceous. MALE. Head dorsally with conspicuous transverse impression behind antennal prominence, antennal cavities separated by minute lamina. Frons with elongate median areola open anteriorly. Eyes moderately large (separated medially above by 1.6 times their radius). Distal joint of maxillary palpi elongate, widening apically, with three blunt glabrous teeth at apical margin. Antennae nearly filiform, from 3rd joint slightly flattened, reaching over half of elytra, with 2nd joint transverse, considerbaly shorter than 3rd, which in its turn slightly shorter than 4th; all antennomeres with relatively short erect pubescence. Pronotum transverse, almost twice as wide as long, trapezoidal, with blunt anterior and acute moderately produced

DIAGNOSIS. In possessing developed sclerotized structures of the internal sack of the aedeagus X. macrolycoides sp.n. is evidently close to X. approximans Bourgeois, of the Central and East Himalayas [Kazantsev, 1992], readily distinguishable by the twisted apical portion of the median piece and other details of the aedeagus (Figs. 5253).


New and little known species of Lycidae (Coleoptera) from China
laterally posterior angles; almost straight anteriorly and slightly bisinuate posteriorly; one third anteriorly and almost same width laterally with coarse punctuation; disk with transverse impression and conspicuous narrow median furrow reaching posterior margin. Scutellum nearly parallel-sided and straight at apex. Elytra long (3 times as long as wide humerally), parallelsided, with almost equally developed, except at apex, primary costa, irregularly granulose in interstices. Pubescence erect, short and dense. Aedeagus relatively long and straight, with free elongate parameres (Figs. 5456). FEMALE. Unknown. Length: 5.0 mm. Width (humerally): 1.4 mm.

263

Literature
Bocak L. 1996. A new species of the genus Platycis from China (Coleoptera: Lycidae) // Folia Heyrovskiana Vol.4. No.1. P.12. Bocakovб M. 1997. Revision of the genus Melaneros from China with a note on Ditoneces (Coleoptera, Lycidae) // Acta Soc. Zool. Bohem. Vol.61. P.175190. Bocakovб M. 2000. Review of the genus Libnetis from China with descriptions of several species from Thailand (Coleoptera: Lycidae) // Acta Soc. Zool. Bohem. Vol.64. P.223234. Kazantsev S. 1988. [To the knowledge of Lycidae (Coleoptera) (On the status of the species Dictyoptera erythroptera Baudi)] // Fauna i ekologiya nasekomykh Vietnama. Moscow: Nauka Publ. P.168170 [in Russian]. Kazantsev S. 1992. To the knowledge of oriental Cantharoidea (Coleoptera). On the Lycidae of Himalaya. Part II // Entom. Basil. Vol.15. P.261266. Kazantsev S. 1996. A review of Pyropterus subgen. Helcophorus Fairmaire with the description of two new oriental species (Lycidae, Coleoptera) // Elytron. Vol.10. P.8188. Kazantsev S. 2000a. New lycids from China (Coleoptera, Lycidae) // Miscel.lania Zoologica Vol.23. No.2. P.7982. Kazantsev S. 2000b. Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China // Elytron Vol.14. P.99109. Kazantsev S. 2001. New species of the genus Melaneros Fairmaire, 1879 from Nepal, China and the USA (Coleoptera, Lycidae) // Russian Entom. J. Vol.10. No.1. P.1316. Kleine R. 1925. Fauna Buruana. Coleoptera, Fam. Lycidae (2. Beitrag zur Kenntnis der Lycidae) // Treubia Bd.7. H.1. S.3137. Pic M. 1921. Contribution а lйtude des Lycides // LEchange, hors texte No.404410. P.128.

DIAGNOSIS. L. taiwanus sp.n. evidently belongs in the same group as L. opacus Pic and L. xilingensis Kazantsev, differing from both by the conspisuous areola on the frons, details of the aedeagus (Figs. 5456), etc. ACKNOWLEDGEMENTS. I wish to express my deep gratitude to Dr. G. Medvedev and Dr. B. Korotyaev (Zoological Institute, St. Petersburg) for enabling me to work with the Lycidae collection and to Mr. Chi-Feng Lee (National Taiwan University, Taipei) for providing interesting material from Taiwan.

Holotype : Taiwan: Kaohsiung, Tengji, 22.VI.2000, C.-F. Lee leg. (SVK).