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Arthropoda Selecta 12 (1): 4765

ї ARTHROPODA SELECTA, 2003

Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaeartic -LEFF= Simon, 1885 Пауки (Arachnida: Aranei) Азербайджана. 2. Обзор пауков-волков (Lycosidae) отмеченных в фауне республики с описанием трех новых видов и краткий обзор пауков рода -LEFF= Simon, 1885 Восточной Палеарктики Yuri M. Marusik1, Elchin F. Guseinov2 & Seppo Koponen Ю.М. Марусик1, Э.Ф. Гусейнов2, С. Копонен3
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Institute for Biological Problems of the North, Portovaya Str. 18, Magadan, 685000 Russia. E-mail: yurmar@mail.ru Институт биологических проблем Севера, ДВО РАН, ул. Портовая 18, Магадан 685000 Россия. Institute of Zoology, block 504, passage 1128, Baku 370073 Azerbaijan. E-mail: elchin-f@artel.net.az Институт зоологии АН Азербайджана, квартал 504, проезд 1128, Баку 370073 Азербайджан. Zoological Museum, University of Turku, FIN-20014 Turku Finland. E-mail: sepkopo@utu.fi

KEY WORDS: Aranei, Lycosidae, Evippa, wolf spiders, check-list, Caucasus, Azerbaijan, new records, new species. КЛЮЧЕВЫЕ СЛОВА: Aranei, Lycosidae, Evippa, пауки-волки, систематический список, Кавказ, Азербайджан, новые находки, новые виды. ABSTRACT: Three species new to science are described from Azerbaijan: Evippa apsheronica sp.n. (), E. caucasica sp.n. () and Pardosa gusarensis sp.n. (). A survey of Evippa Simon, 1882 reported from the former USSR is given and six species are illustrated. Evippa potanini Schenkel, 1963 syn.n. was found to be a junior synonym of E. sjostedti Schenkel, 1936. Taxonomic status of Allohogna Roewer, 1955 and generic placement of some species are discussed. Trochosa hispanica Simon, 1870 was found in Azerbaijan and Caucasus for the first time. According to literature data and material studied the check-list of Azerbaijanian lycosids encompass 69 species belonging to 14 genera: Allohogna (1), Alopecosa (14), Arctosa (4), Aulonia (2), Evippa (2), Geolycosa (1), Hogna (4), Lycosa (1), Mustelicosa (1), Pardosa (29), Pirata (3),Trochosa (4),Wadicosa (1),Xerolycosa (2). Records of 12 species and one genus are doubtful and require confirmation. РЕЗЮМЕ: Описаны три новых вида из Азербайджана: Evippa apsheronica sp.n. (), E. caucasica sp.n. () и Pardosa gusarensis sp.n. (). Приведен обзор видов рода Evippa Simon, 1882 зарегистрированных в СССР и проиллюстрированы 6 видов. Обнаружено, что Evippa potanini Schenkel, 1963 syn.n. является младшим синонимом E. sjostedti Schenkel, 1936. Обсуждается таксономический статус рода Allohogna Roewer, 1955 и родовая принадлежность некоторых видов. Trochosa hispanica Simon, 1870 обнаружена впервые в фауне Азербайджана и всего Кавказа. Согласно литературным и новым данным фауна пауков-волков республики включает 69 видов относимых к 14 родам: Allohogna (1), Alopecosa (14), Arctosa (4), Aulonia (2), Evippa (2), Geolycosa (1), Hogna (4), Lycosa (1), Mustelicosa (1), Pardosa (29), Pirata (3), Trochosa (4), Wadicosa (1), Xerolycosa (2). Находки 12 видов и одного рода являются сомнительными и нуждаются в подтверждении.

Introduction
Spiders of the family Lycosidae are rather well studied in Caucasus. The first species of wolf spiders were reported and described from Azerbaijan by L. Koch [1878] in his survey of Caucasian arachnids. Later, Schmidt [1895] added a few more species for Caucasus and Azerbaijan. A detailed survey of Caucasian spiders was provided by Verzhbitsky [1902]. He listed 147 species and 3 subspecies. According to the latest data [Marusik & Guseinov, 2003] 83 species of wolf spiders belonging to 12 genera were known to occur in Caucasus, of them 59 species belonging to 12 genera were known from Azerbaijan. Two additional genera Evippa and Wadicosa were added by us [Marusik & Guseinov, 2003] to Azerbaijan and Caucasian faunas. By the number of species known in Azerbaijan wolf spiders are the third largest family. Higher species diversity was reported for Linyphiidae [89 species, cf.


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen

Mikhailov, 2002] and Salticidae [82 species, cf. Logunov & Guseinov, 2001]. Only one paper was devoted specifically to lycosids from Azerbaijan [Zyuzin & Logunov, 2000]. In the beginning of this project, the main goal of our paper was to survey all literature data about wolf spiders of Azerbaijan and to give comments on the status of species and genera found there. However the scope of our study was extended when, in checking some old and new material, we found three new species belonging to Evippa and Pardosa. Because Evippa is a poorly studied genus in east Palaearctic we decided to provide a brief illustrated review of species known in the former Soviet Union. While it was found that several species records from Azerbaijan are doubtful we are not removing these species from the country-list until more material is studied and necessary revisions done. Lycosa felina L. Koch, 1878, mentioned from Azerbaijan [cf. Mikhailov, 1997], is not listed here. It was described from Georgia (Azkhur) and was mistakenly assigned to Azerbaijan.

Descriptions of new species and survey of Evippa reported from the former Soviet Union
Evippa Simon, 1882
Thirty-four species are known to belong to this genus. Evippa species occur in south Palaearctic, southern China, northern parts of India and Africa south to the Congo River. Ten species of this genus were listed from the former USSR [Mikhailov, 1997]. All of them, except for E. eltonica Dunin, 1994 were known from the Asian part of the former USSR. Until recently this genus was unknown from Azerbaijan and the Caucasus as a whole. The nearest records of Evippa lie in northern Ciscaspia (Elton Lake) and Turkmenistan. Thanks to the courtesy of Drs D.V. Logunov & A.A. Zyuzin we were able to examine all central Asian Evippa. Taking this opportunity we give figures of these species, but without text, which will be provided in a special revision.

Evippa kronebergi Roewer, 1955
Lycosa aculeata Kroneberg, 1875: 38, pl. 5, fig. 46ac (). Evippa aculeata: Simon, 1895: 342; Mikhailov, 1997: 124. Pardosa a.: Charitonov, 1932: 42 Acantholycosa a.: Reimoser, 1935: 171; Bonnet, 1955: 132. Evippa kronebergi Roewer, 1955: 154 (replacement name, by thought that Lycosa aculeata preoccupied by Sundevall, 1833); Mikhailov, 1998: 19; Platnick, 2002.

Material and methods
All L. Kochs [1878] records were made around Baku and here we list these records as Apsheron Peninsula. Below we list geographical names which are spelled in more than one way: Absheron = Apsheron; Kusary = Gusar; Nakhchyvan = Nakhitchevan; Zakataly = Zagatala. The city presently known as Gyandzha, had the name Elisabetpol = Elisawetpol early in the 19th century to beginning of 20th. Later, in Soviet times (mid 1930s) its name was changed to Kirovabad, and in the 1990s its name was name was changed again to the Gyandzha (name used before 19th century). Illustrations were made using both reflected and transmitted light microscopes with drawing devices. Microphotographs were made with a Jeol JSM-5200 SEM in the Zoological Museum, University of Turku. The following abbreviations have been used for collections and museums: CAS Californian Academy of Sciences, San-Francisco; IZBA Institute of Zoology, Baku; YMT Yuri M. Marusiks temporary collection in Zoological Museum, University of Turku; ZMMU Zoological Museum, Moscow University; ZMUT Zoological Museum, University of Turku. Collectors names are also abbreviated: EG Elchin F. Guseinov, YM Yuri M. Marusik. Some standard abbreviations have been used for eyes and their interdistances: AME anterior median eyes; ALE anterior lateral eyes; PME posterior median eyes; AMEPME distance between AME & PME, ш diameter.

Study of all catalogues reveals that Lycosa aculeata Sundevall, mentioned by Roewer [1955] was never described. Sundevall [1833] just listed in his paper Lycosa aculeata (Clerck, 1757) (= Alopecosa aculeata). According to Bonnet [1955] this combination (Lycosa a.) was never used by other authors, and at the time of Kronebergs description his species name was not a homonym of the Clercks name. However Roewers [1955] replacement name is valid, because it was made before 1960 (cf. Article 59.2. of ICZN, 4th edition). Unfortunately the type of this species was lost. Figure 46c provided by Kroneberg [1875] for epigyne resembles those of E. schenkeli ternbergs, 1979 and another species belonging to sjostedti group.

Evippa badchysica ternbergs, 1979 Figs. 2324.
COMMENTS. Known so far only from the type locality, southwestern Turkmenistan.
Evippa badchysica ternbergs, 1979: 65, fig. 1 ().

Evippa beshkentica Andreeva, 1976 nomen nudum [cf. Mikhailov, 1997]. Illustrations and list of material were given, but there was no formal description.

Figs. 112. Copulatory organs of Evippa apsheronica sp.n. (13), E. caucasica sp.n. (46), E. eltonica Dunin (79) and E. sibirica Marusik (1012). 12 epigyne, ventral view, before and after maceration respectively; 3 epigyne, dorsal view; 4, 7, 10 male palp, ventral view; 5, 11 part of tegulum with tegular apophysis and conductor, retrolateral view; 6, 9, 12 embolic complex; 8 tegular apophysis and conductor, retrolateral view. Scale = 0.1 mm. Рис. 112. Копулятивные органы Evippa apsheronica sp.n. (13), E. caucasica sp.n. (46), E. eltonica Dunin (79) и E. sibirica Marusik (1012). 12 эпигина, вид снизу, перед и после мацерации; 3 эпигина, вид сверху; 4, 7, 10 пальпа самца, вид снизу; 5, 11 часть тегулюма с тегулярным отростком и кондуктором, вид сбоку-сзади; 6, 9, 12 эмболюсный отдел; 8 тегулярный отросток и кондуктор, вид сбоку-сзади. Масштаб 0,1 мм.


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen Evippa concolor (Kroneberg, 1875)
Lycosa concolor Kroneberg, 1875: 38, pl. 4, fig. 26ac (). Evippa c.: Simon, 1895: 342.

COMMENTS. This species is distributed from northeastern Kazakhstan to Tuva and Gobi-Altai Aimak in Mongolia [Marusik et al., 2000].

COMMENTS. This species was described from Tajikistan. Its type was lost (Mikhailov, personal communication). Judging from Kronebergs figure of the male palp, showing the tegular apophysis placed medially as in Pardosini, it seems that this species was erroneously assigned to Evippa by Simon [1895].

Evippa sjostedti Schenkel, 1936 Figs. 3034.
Evippa sjцstedti Schenkel, 1936: 304, fig. 106 (). E. potanini Schenkel, 1963: 387, fig. 224 (). Syn.n. Xerolycosa brunneopicta Loksa, 1965: 16, fig. 24 (). Material examined: Holotype of Evippa sjostedti Schenkel, 1936 S.Mong. C.26, 9.V.[19]27; Sven Hedins Exp. Ctr. Asien; S.Mongolie 1927 in Stockhom; holotype Evippa potanini Schenkel, 1963, from Bergkette Bann-Tsagan, Sichuan in NMNH with label Potanin 68, #68, carapace with right legs dissected, no epigyne, vial contains small glass balls, somebody examined the holotype. MONGOLIA: 19 19 1 juv. (YMT), Omnogov (=South Gobi) Aimak, Bayandalai Somon, Zoolen uul (Mt. Range), 43њ21N, 103њ11E, 1700 m, 2730.05.1997 (YM); 6 3 (YMT), Omnogov Aimak, Gurvantes Somon, NW part of Sevren Mt. Range, 43њ38N, 101њ08E, 1900 m, 01.06.1997; 4 15 (YMT), Omnogov Aimak, Noyon Somon, Noyon uul (Mt. Range), 1900 m, 3031.05.1997 (YM); 2 (YMT), Bayanhkongor Aimak, Bayanlig Somon, Ikh-Bogd Mt. Range, south foothils, 1600 m, 04.06.1997 (YM); 24 1 juv., Bayanhkongor Aimak, Bayanlig Somon, Bor-Tolgoi, 44њ06N 100њ56E, 1400 m, 0204.06.1997 (YM); 7 1 juv. (ZMUT) [09] Bayanhkongor Aimak, Bogd Somon, Ikh-Bogd Mt. Range, IkhBogd Pass, 44њ43N, 100њ52E, 20002100 m, 04.06.1997 (YM); 1 (YMT), Bayanhkongor Aimak, Bogd Somon, Ikh-Bogd Mt. Range, north foothils, 1600 m, 04.06.1997 (YM).

Evippa eltonica Dunin, 1994 Figs. 79, 1315.
Evippa eltonica Dunin, 1994: 243, fig. 14 (). Evippa sp.: Eskov & Marusik, 1995: 65, fig. 63 ().

COMMENTS. Besides the type locality, Elton Lake, at the eastern border of Volgograd Area, it was found in adjacent northwestern Kazakhstan, in environs of Dzhanybek [Eskov & Marusik, 1995].

Evippa onager Simon, 1895
Evippa onager Simon, 1895c: 341 (). E. o.: ternbergs, 1979: 67, fig. 1 ().

COMMENTS. This species was described from China (Nan-Shan-Kou, Xinjiang (43?171N, 93.786?E), while listed by Platnick [2002] as distributed in Turkmenistan. ternbergs [1979] had not studied the types of this species and his identification was clearly mistaken. Judging from the distribution of E. sjostedti Schenkel, 1936, E. onager could be a senior synonym of it. In Figs. 1922, 2829 we show what species was treated as E. onager by ternbergs.

Evippa potanini Schenkel, 1963 see E. sjostdedti. Evippa praelongipes (O. Pickard-Cambridge, 1870)
COMMENTS. Study of numerous specimens identified from Central Asia by ternbergs as E. praelongipes (O. Pickard-Cambridge, 1870) reveals that one specimen labeled as E. praelongipes was, in fact, a syntype of E. turkmenica ternbergs, 1979. It seems that E. praelongipes was reported from Central Asia (environs of Krasnovodsk) by Schmidt [1895]. Schmidt even described a female of this species for the first time. Judging from his description and Zyuzin (personal communication), Schmidt dealt with E. turkmenica. This species was also reported from the territory of the former USSR by Spassky [1952] and by Ponomarev [1988]. Both these publications are lacking figures, exact localities and material examined. Most probably this species does not occur in the former USSR.
Evippa p. Alderweireldt, 1991: 369, fig. 5.15 ().

COMMENTS. Study of the types of E. sjostedti and E. potanini as well as numerous Mongolian specimens leaves no doubt that the two names are synonyms. Figures of E. sjostedti in Loksa [1965, fig. 29] and Izmailova [1980, fig. 1] correspond to Pardosa adustella Roewer, 1951. There were no specimens of this species among material from Central Asia studied so this species has to be excluded from the fauna of the former USSR.

Evippa turkmenica ternbergs, 1979 Fig. 35.
COMMENTS. The species description was based on two adult females and two juveniles. ternbergs did not designate a holotype. Study of the type specimens of Evippa described by ternbergs, reveals no types of E. turkmenica, but instead of them we found one vial with a species label E. praelongipes (O. Pickard-Cambridge, 1870) and a geographical label the same as one syntype of E. turkmenica (Sandy-Kachi). It seems that type series of this species was not labeled as a whole. Of these 12 species reported from the former Soviet Union three were misidentified, E. onager, E. praelongipes and E. sjostedti (=E. potanini), and one species name is invalid (nomen nudum) and therefore the known fauna of the former USSR comprises only 7 correctly identified species. East Palaearctic Evippa can be attributed to two clear species groups: the sjostedti-group (sjostedti, turkmenica, helanshangensis Peng, Yin & Kim, 1996, a few undescribed species and possibly onager and benevola (O. Pickard-Cambridge, 1885), and the eltonica-group (eltonica, sibirica, fujianensis Peng, Yin & Kim, 1996, onager sensu ternbergs and some undescribed species). The position of other east Palaearctic species including E. badchysica and E. schenkeli remains
Evippa schenkeli ternbergs, 1979: 67, fig. 1 ().

Evippa schenkeli ternbergs, 1979 Figs. 2527.
COMMENTS. Known so far only from the type locality, southwestern Turkmenistan.
Evippa schenkeli Sternbergs, 1979: 65, fig. 1 ().

Evippa sibirica Marusik, 1995 Figs. 1012.
Evippa sibirica Marusik, in Eskov & Marusik, 1995: 64, fig. 58 62 ().


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Figs. 1327. Epigynes and body of Central Asian Evippa: E. eltonica Dunin (1315), E. sibirica Marusik (1618), E. onager Simon sensu ternbergs (1979) (1922), E. badchysica ternbergs (2324, holotype) and E. schenkeli ternbergs (2527, holotype). 13 14, 1617, 1921, 23, 27 epigyne, ventral view; 15, 18, 22, 24, 27 epigyne, dorsal view; 25 body. 1315 specimens from Dzanybek, NW Kazakhstan; 1618 paratypes from NE Kazakhstan, 19 from Repetek; 2022 from Morgunovka. 1314, 16 17 and 1921 showing epigynal variations. Scale for all figures except for 25 = 0.1. Рис. 1327. Эпигина и внешний вид среднеазиатских видов Evippa: E. eltonica Dunin (1315), E. sibirica Marusik (1618), E. onager Simon sensu ternbergs (1979) (1922), E. badchysica ternbergs (2324, голотип) и E. schenkeli ternbergs (2527, голотип). 1314, 1617, 1921, 23, 27 эпигина, вид снизу; 15, 18, 22, 24, 27 эпигина, вид сверху; 25 внешний вид. 1315 экземпляры из Джаныбека, СЗ Казахстан; 1618 паратипы из СВ Казахстана, 19 из Репетека; 2022 из Моргуновки. 1314, 1617 и 1921 вариации эпигины. Масштаб для всех рисунков кроме 25 0,1мм.


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen
long light hairs. Palp as in Figs. 46. Cymbium dark brown, other joints light brown, upper part of cymbium with 6 macrosetae, embolus round and, as in other congeners, most of it hidden by tegulum, tegular apophysis with bill-shaped outgrowth closely attached to conductor. Leg joint length femur patella tibia metatarsus tarsus total I 2.30 1.00 1.83 2.00 0.88 8.01 II 2.38 0.95 1.85 1.90 1.03 8.11 III 2.25 1.03 1.70 2.00 1.10 8.08 IV 2.75 1.10 2.25 3.13 1.25 10.48 Spination of legs: I: femur 3d+2p+2r, patella 2d+1p+1r, tibia 5v+2d+2p, metatarsus 3v+1p; II: femur 3d+2p+2r, patella 2d+1p+1r, tibia 5v+2d+2p, metatarsus 3v+1p; III: femur 3d+2p+2r, patella 2d+1p+1r, tibia 2v+2d+2p+2r, metatarsus 2v+3p+2r; IV: femur 3d+2p+2r, patella 2d+1p+1r, tibia 2v+2d+2p+2r, metatarsus 2+1v+3p+2r. Female unknown. DIAGNOSIS. This species belongs to the eltonica species group. It can be easily separated from all known species of this group by the shape of tegular apophysis, in which the bill shaped outgrowth has a straight horizontal lower margin (rounded in E. eltonica and E. sibirica). COMMENTS. The two species found in Azerbaijan each known from one sex and can not be conspecific because they belong to different species groups.

unclear. The two above mentioned groups can be easily separated by the shape of tegular apophysis, embolic complex and epigyne from the type species of the genus (Lycosa arenaria Audouin, 1827) and possibly represent a separate taxon.

Species descriptions
Evippa apsheronica sp.n. Figs. 13.
Material: Holotype (ZMMU), central-eastern Azerbaijan, Apsheron Peninsula, Baku, near Ganly-Gyol Lake, 40њ21.46N, 49њ48.36E, 11.05.1999 (EG). Paratypes: 3 & 1 juv. (ZMMU and YMT), same locality, 06.06.2003 (EG & YM).

DESCRIPTION. Female. Body 10.0 long. Carapace: 4.0 long, 3.1 wide, light brownish, with light median band, eye field darkened. Sternum light-grey with yellow median band. Labium dark-grey. Gnathocoxae and chelicerae yellow. Legs yellow with light-grey rings, tibia with two light and two dark rings. Abdomen dorsally light brownish with pattern of yellow spots of different sizes, heart mark dark, posterior half with large transverse spot, venter light-yellow. Leg joint length Leg femur patella tibia metatarsus tarsus total I 3.25 1.60 2.45 2.63 1.70 11.63 II 3.13 1.63 2.25 2.50 1.70 11.21 III 3.13 1.50 2.15 3.13 1.75 11.66 IV 4.25 1.75 3.00 4.25 2.13 15.38 Spination of legs: I: femur 3d+1p+2r, patella 2d+1p+1r, tibia 2v+2d+2p+2r, metatarsus 2v+2p; II: femur 3d+2p+2r, patella 2d+1p+1r, tibia 1+1v+2p+2r, metatarsus 2v+2p; III: femur 3d+2p+2r, patella 2d+1p+1r, tibia 1+1v+2d+2p+2r, metatarsus 2v+2d+2p+2r; IV: femur 3d+2p+1r, patella 2d+1p+1r, tibia 2v+2d+2p+2r, metatarsus 2+1v+2d+2p+2r. Epigyne as in Figs. 13, with almost rectangular septum and round fovea with indistinct upper margin, endogyne with accessorial glands. Male unknown. DIAGNOSIS. In the shape of its epigyne the new species resembles the Central Asian E. schenkeli ternbergs, 1979 (cf. Fig. 26) and the Algerian E. jocquei Alderweireldt, 1991 (cf. Fig. 8.3 in Alderweireldt [1991]), but can be easily separated from them by its more rectangular septum and round fovea. From all Evippa species E. apsheronica sp.n. can be easily distinguished by having accessorial glands.

ity.

ETYMOLOGY. The species is named after its type local-

Pardosa gusarensis sp.n. Figs. 3638.
Material: Holotype (ZMMU), Azerbaijan, Gusar Distr., foothill of Bazar-Dyuzyu Mt., 3000 m, 09.08.2001 (EG).

Evippa caucasica sp.n. Figs. 46.
Material: Holotype (ZMMU), central-eastern Azerbaijan, Gobustan, Beyuk-dash hill, 40њ05N, 49њ25E, 15.04.2001 (EG). Paratype , central-eastern Azerbaijan, ca 75 km N of Baku, W of Kylyazi Vill., 40њ51.5N, 49њ11.5E, 260 m, semi-desert, river bank, 07.06.2003 (YM).

ETYMOLOGY. The species epithet refers to the type locality. DESCRIPTION. Male. Body 6.3 long. Carapace: 3.25 long, 2.4 wide, dark gray-brown, eye field covered with dense light hairs. Sternum and labium dark brown. Gnathocoxae grey, apical portion colored as labium. Chelicerae dark brown. Legs grey-brown with yellow rings, tarsi yellow. Abdomen grey, without pattern, sides and venter covered with dense and

ETYMOLOGY. The specific epithet derived from the type locality. DESCRIPTION. Body 7.15 long. Carapace: 3.38 long, 2.50 wide, dark brown with sparse light hairs, eye-field and margins black. Sternum dark brown, labium and gnathocoxae yellow. Chelicerae dark brown with yellow spots. Legs dark grey-brown with yellow spots, tarsi yellow. Abdomen dark grey with lighter venter. Epigyne as in Figs. 3638, with two separate large apical pockets, long and thin septal stem, fovea absent. Septum looks as if fused with epigynal plate, its margins unclear. Leg joint length femur patella tibia metatarsus tarsus total I 2.50 1.30 2.38 2.13 1.13 9.44 II 2.50 1.25 2.20 2.15 1.00 9.1 III 2.45 1.15 2.00 2.65 1.13 9.38 IV 3.15 1.25 3.00 2.80 1.75 11.95 Spination of legs: I: femur 3d+3p+2r, patella 2d+1p+1r, tibia 3v+2d+2p+2r, metatarsus 2v+1p+1r; II:femur 3d+2p+2r, patella 2d+1p+1r, tibia 3v+2d+2p+2r, metatarsus 2v+1p+1r; III: femur 3d+2p+2r, patella 2d+1p+1r, tibia 2v+2d+2p+2r, metatarsus 2v+2p+2r; IV: femur 3d+2p+1r, patella 2d+1p+1r, tibia 2v+2d+2p+2r, metatarsus 2v+2p+2r. DIAGNOSIS. This species is closely related to another high-alpine Caucasian species, P. ibex Buchar & Thaler, 1998. P. gusarensis sp.n. has larger and more widely spaced apical pockets, their width exceeds width of septal stem (width of stem and pockets equal in P. ibex). Upper margin of apical pockets in P. gusarensis sp.n. are almost horizontal, while in sibling species are directed upward. By the shape of septum which appears to be fused with epigynal plate these species can be easily separated from all other Pardosa species known to us.


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Figs. 2835. Copulatory organs of Evippa onager Simon sensu ternbergs (1979) (2829), E. sjostedti Schenkel (3034) and E. turkmenica ternbergs (35, ?syntype). 28, 30 male palp, ventral view; 29 male palp, retrolateral view; 31 part of tegulum with tegular apophysis and conductor; 32 embolic complex; 3334 epigyne ventral view. 2829 specimen from Turkmenia; 33 holotype of E. sjostedti; 3032 & 34 specimens from Mongolia. Scale = 0.1 mm. Рис. 2835. Копулятивные органы Evippa onager Simon sensu ternbergs (1979) (2829), E. sjostedti Schenkel (3034) и E. turkmenica ternbergs (35, синтип). 28, 30 пальпа самца, вид снизу; 29 пальпа самца, вид сбоку-сзади; 31 часть тегулюма с тегулярным отростком и кондуктором; 32 эмболюсный отдел; 3334 эпигина, вид снизу. 2829 экземпляры из Туркмении; 33 голотип E. sjostedti; 3032 & 34 экземпляры из Монголии. Масштаб 0,1 мм.

COMMENTS. Endogynes of two related species are unique in having closely separated receptacula converging basally. Buchar & Thaler [1998] placed this species among proxima- and wagleri-groups of Pardosa. Judging from the rather peculiar shape of its epigyne this species may form a separate species group.

Pardosa sp.

Figs. 3941, 51.

Material examined: 3 (ZMUT), Azerbaijan, Gusar Distr., foothill of Bazar-Dyuzyu Mt., 3000 m, 41.20њN, 47.89њE, 09.08.2001 (EG).


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen

Figs. view. 37 Рис. 37 & 40

3641. Epigyne of Pardosa gusarensis sp.n. (3638) and Pardosa sp. (3941). 3637, 39-40 ventral view; 38, 41 dorsal & 40 view after maceration. Scale = 0.1 mm. 3641. Эпигина Pardosa gusarensis sp.n. (3638) и Pardosa sp. (3941). 3637, 3940 вид снизу; 38, 41 вид сверху. после мацерации. Масштаб 0,1 мм.

COMMENTS. Specific placement of our specimens is unclear. Shape of epigyne and receptacula indicate close relationships with P. aquila Buchar & Thaler, 1998 and P. italica Tongiorgi, 1966. Lack of males and comparative material of the two sibling species does not allow us to attribute our specimens to a known species.

Survey of lycosid species reported from Azerbaijan
Allohogna Roewer, 1955 Allohogna singoriensis (Laxmann, 1770) Figs. 4246.
Lycosa s.: Fuhn & Niculescu-Burlacu, 1971: 198, fig. 96ae ().

RECORDS: Apsheron Pen. [sub Lycosa s. L. Koch, 1878; Dunin, 1984], Muganskaya steppe [Dunin & Mamedov, 1992]. COMMENTS. In this paper we decided to provide figures of A. singoriensis, the type species of the genus for two reasons. First, there are few diagnostic drawings of it. The other reason is to show that Allohogna is distantly related to Lycosa Latreille, 1804 and can not be treated as its junior synonym [cf. Platnick, 2002]. Lycosa is rather obscure genus,

because its type species Aranea tarantula Rossi, 1790 is known only by figures made in the 19th century. Our judgment about Lycosa is based on the recent definition of the genus given by Zyuzin & Logunov [2000]. Zyuzin (personal communication) studied all available types of Lycosa and provided [Zyuzin & Logunov, 2000] figures of two species L. praegrandis C.L. Koch, 1836 and L. narbonensis Walckenaer, 1806. However, the generotype of this genus remains unredescribed and only the eyes of L. tarantula were illustrated [fig. 1 in Zyuzin, 1985]. Unlike in Allohogna, the cymbium in Lycosa is assymetrical, its tegular apophysis is very massive, subtegulum turned to prolateral side. Females of Lycosa have no apical pockets in epigyne and its septum longer than wide, apical part of epigynal plate with anterior elevation (in Allohogna: 2 widely spaced pockets, septum anchor shaped, its length subequal to width, anterior elevation absent). L. tarantula and A. singoriensis can be easily separated by eye formula [cf. figs. 1 & 3 in Zyuzin, 1985]: anterior row of eyes straight in A. singoriensis (recurved in L. tarantula), lateral margins of ALE spaced equally with PME (ALE-ALE < PME-PME); AME-PME < ш of AME (AME-PME = ш of AME). To us, Allohogna is clearly different from Lycosa and has to be removed from synonymy. COMMENTS. According to Platnick [2002] this species has a Palaearctic distribution but its actual range is possibly much smaller [cf. Marusik et al., 2000].


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Figs. 4250. Copulatory organs of Allohogna singoriensis (Laxmann) (4246) and Pardosa tatarica (Thorell) (4750). 42 palp, dorsal view; 4344 bulbus, ventral view and view from above respectively; 4546 epigyne, ventral and dorsal view respectively; 47, 49 apical portion of the bulbus, ventro-apical view; 48, 50 terminal portion of the bulbus, view from above. 4246 specimens from Tuva, 4748 from Azerbaijan, 49-50 from Greece. Scale = 0.1 mm if not otherwise indicated. Рис. 4250. Копулятивные органы Allohogna singoriensis (Laxmann) (4246) и Pardosa tatarica (Thorell) (4750). 42 пальпа, вид сверху; 4344 бульбус, вид снизу и сверху; 4546 эпигина, вид снизу и сверху; 47, 49 верхняя часть бульбус; 48, 50 верхняя часть бульбуса, вид сверху. 4246 экземпляры из Тувы, 4748 из Азербайджана, 4950 из Греции. Масштаб 0,1 мм, если не указан другой.

Alopecosa Simon, 1885 Alopecosa accentuata (Latreille, 1817)
Tarentula a.: Holm, 1947: 18, pl. 3, fig. 2324, pl. 9, fig. 14 (). Alopecosa a.: Lugetti & Tongiorgi, 1969: 13, fig. 4af (). A. a.: Fuhn & Niculescu-Burlacu, 1971: 142, fig. 65ae (). A. a.: Heimer & Nentwig, 1991: 314, fig. 845 (). A. a.: Roberts, 1998: 241, fig. (). Material examined: 1 , Lenkoran Distr., Avrora, 23.04.2001 (EG).

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [sub Tarentula a., Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992]. COMMENTS. This species is treated by Platnick [2002] as a Palaearctic form, and listed by Mikhailov [1997] as a species distributed through the whole former Soviet Union. Study of numerous specimens from Siberia and Far East has not revealed any individuals of this species.

Alopecosa aculeata (Clerck, 1757)
Tarentula a.: Holm, 1947: 19, pl. 3, fig. 32, pl. 9, fig. 1718 ().


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen
RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [sub Tarentula c., Dunin, 1989]. COMMENTS. Known through the whole Europe and Central Asia eastward to Tibet [cf. Song et al., 1999; Platnick, 2002].

Alopecosa a.: Lugetti & Tongiorgi, 1969: 25, fig. 7af (). A.a.: Fuhn & Niculescu-Burlacu, 1971: 144, fig. 66ae (). A.a.: Kronestedt, 1990: 204, fig. 1AB, 3A, 4AB, 5A, a, 6A, C, E, 7A, 8AD, 10A, 11A, 12A, 13AE (). A.a.: Heimer & Nentwig, 1991: 312, fig. 843 (). A. a.: Roberts, 1998: 239, fig. ().

RECORDS: Sheki-Zagatala area [sub Tarentula a., Dunin, 1989] COMMENTS. This species has circum-Holarctic range [Marusik et al., 2000].

Alopecosa etrusca Lugetti et Tongiorgi, 1969
RECORDS: (sub.Tarentula e.): Lenkoran area [Guseinov, 1999]. COMMENTS. This species so far is known only from Italy [Platnick, 2002], and most probably records from Azerbaijan refer to other species (eg. A. cursor).
Alopecosa e. Lugetti & Tongiorgi, 1969: 58, fig. 16af ().

Alopecosa albofasciata (Brullй, 1832)
Alopecosa a.: Lugetti & Tongiorgi, 1969: 43, fig. 12af (). A. a.: Fuhn & Niculescu-Burlacu, 1971: 145, fig. 67af (). Material examined: 14 17 5 juv., SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 23 28.04.2001 (YM & EG).

Alopecosa pentheri (Nosek, 1905)
Alopecosa cursor p.: Lugetti & Tongiorgi, 1969: 55, fig. 15ac (). A. insignis: Lugetti & Tongiorgi, 1969: 57, fig. 15de (, elevated from subspecies of A. cursor). A. p.: Thaler et al., 2000: 1073, fig. 34, 42, 4647 ().

RECORDS: (sub Tarentula a.): Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999] COMMENTS. A. albofasciata known from Spain to Central Asia [Thaler et al., 2000; Platnick, 2002].

Alopecosa alpicola (Simon, 1876)
RECORDS: (sub Tarentula a.): Lenkoran area [Guseinov, 1999]. COMMENTS. Listed by Platnick [2002] as a Palaearctic species. We think all records in Asia refer to similar species. Without doubt records from Azerbaijan refer to another species.
Alopecosa a.: Lugetti & Tongiorgi, 1969: 70, fig. 20af ().

RECORDS: Apsheron Pen. [Dunin, 1984], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [sub Tarentula p., Guseinov, 1999]. COMMENTS. Known from Greece and Yugoslavia (Thaler et al., 2000) and from Bulgaria to Azerbaijan [Platnick, 2002].

Alopecosa pulverulenta (Clerck, 1757)
Alopecosa p.: Kronestedt, 1990: 217, fig. 4F, 5D, 8HI, 10D, 11D, 12D, 13KM (). A. p.: Roberts, 1995: 224, fig. (). A. p.: Roberts, 1998: 238, fig. ().

Alopecosa cronebergi (Thorell, 1875)
RECORDS: (sub Tarentula c.): Lenkoran area [Guseinov, 1999]. COMMENTS. This species is known from two females described from Derbent (Daghestan, Russia) and was never illustrated, and therefore the record from Azerbaijan may refer to another species.

RECORDS: (sub Arctosa strandi): Sheki-Zagatala area [Dunin, 1989]. COMMENTS. This species has a trans-Palaearctic range [Marusik et al., 2000].

Alopecosa schmidti (Hahn, 1835)
Alopecosa s.: Lugetti & Tongiorgi, 1969: 86, fig. 25ag (). A. s.: Fuhn & Niculescu-Burlacu, 1971: 160, fig. 77ag (). A. s.: Heimer & Nentwig, 1991: 312, fig. 840 ().

Alopecosa cuneata (Clerck, 1757)
Alopecosa c.: Lugetti & Tongiorgi, 1969: 33, fig. 9ag (). A. c.: Fuhn & Niculescu-Burlacu, 1971: 148, fig. 69af (). A. c.: Roberts, 1985: 140, fig. 61d (). A. c.: Kronestedt, 1990: 217, fig. 4E, 5C, 8G, 10C, 11C, 12C, 13HI (). A. c.: Heimer & Nentwig, 1991: 312, fig. 841 (). A. c.: Roberts, 1995: 224, fig. (). A. c.: Roberts, 1998: 238, fig. (). Material examined: 1 (YMT), SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001 (YM)

RECORDS: (sub Tarentula s.): Sheki-Zagatala area [Dunin, 1989]. COMMENTS. Exact range is unclear, all records at least from Siberia are based on missidentified specimens.

Alopecosa striatipes (C. L. Koch, 1839)
Alopecosa s.: Lugetti & Tongiorgi, 1969: 79, fig. 22af (). A. s.: Fuhn & Niculescu-Burlacu, 1971: 164, fig. 80af (). A. s.: Heimer & Nentwig, 1991: 312, fig. 838 (). A. s.: Roberts, 1998: 243, fig. (). Material examined: 1 (YMT), Apsheron Pen., Kergez Hill, 28.04.2000 (EG).

RECORDS: (sub Tarentula c.): Lenkoran area [Guseinov, 1999]. COMMENTS. This species has a trans-Palaearctic range [Marusik et al., 2000].

Alopecosa cursor (Hahn, 1831)
Alopecosa c.: Lugetti & Tongiorgi, 1969: 52, fig. 14ag (). A. c.: Fuhn & Niculescu-Burlacu, 1971: 150, fig. 70ae (). A. c.: Heimer & Nentwig, 1991: 314, fig. 847 (). A. c.: Roberts, 1998: 243, fig. (). Material examined: 1 (YMT), Apsheron Pen., env. of Gyurgyan Vill., 40њ24N, 50њ16E, semidesert, 17.04.2001 (YM).

RECORDS: Apsheron Pen. [Dunin, 1984], Lenkoran area [sub Tarentula s., Guseinov, 1999]. COMMENTS. Exact range of this species is unclear, it seems that it is distributed from Europe to Central Asia.

Alopecosa sulzeri (Pavesi, 1873)
Alopecosa s.: Lugetti & Tongiorgi, 1969: 67, fig. 19ae (). A. s.: Fuhn & Niculescu-Burlacu, 1971: 167, fig. 81ae ().


Spiders (Arachnida, Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae)
RECORDS: Apsheron Pen. [Dunin, 1984]. COMMENTS. This species is treated as Palaearctic species by Platnick [2002]. We think records from Azerbaijan and whole former Soviet Union and China may refer to other species.
A. s.: Heimer & Nentwig, 1991: 312, fig. 835 ().

57

Arctosa tbilisiensis Mcheidze, 1946
Arctosa t.: Thaler et A. t.: Alderweireldt, Material examined: Distr., env. of Aurora (YM). al., 2000: 1076, 2002: 183, fig. 1 juv (YMT), Vill., 38њ40N, fig. 1516, 4951. 15 (). SE Azerbaijan, Lenkoran 48њ52E, 2328.04.2001

Alopecosa taeniopus (Kulczyсski, 1895)
Alopecosa t.: Lugetti & Tongiorgi, 1969: 84, fig. 24ad (). A. t.: Fuhn & Niculescu-Burlacu, 1971: 168, fig. 82ae ().

RECORDS: (sub Tarentula): Lenkoran area [Guseinov, 1999]. COMMENTS. Treated by Platnick [2002] as distributed from Bulagaria to China, while we think that records from Central Asia and China may refer to other species.

Arctosa C.L. Koch, 1847 Arctosa cinerea (Fabricius, 1777)
Arctosa c.: Lugetti & Tongiorgi, 1965: 171, fig. I.14 (). A. c. Fuhn & Niculescu-Burlacu, 1971: 176, fig. 86ae (). A. c. Roberts, 1985: 148, fig. 64d (). A. c. Heimer & Nentwig, 1991: 320, fig. 852 (). A. c. Roberts, 1995: 229, fig. (). A. c. Roberts, 1998: 247, fig. ().

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999], Geokchai Distr. (Geokchai River) [Alderweireldt, 2002]. COMMENTS. Known from Macedonia to Caucasus (Thaler et al., 2000). Brief comparison of specimens from Lenkoran with these from more northern populations gave some evidence that they may belong to different species. It is worth mentioning that this species, judging from its general appearance (bright coloration in contrast to cryptic pattern of Arctosa, shiny carapace) and copulatory organs are clearly not congeneric with the generotype A. cinerea (Fabricius, 1777) (cf. above mentioned figures with fig. I.14 in Lugetti & Tongiorgi [1965] and fig. 48 in Thaler et al. [2000]). Zyuzin [1985] was probably the first to point out that A. tbilisiensis was not a true Arctosa (not congeneric with A. cinerea). His conclusion was made on the basis of study of eye formula.

RECORDS: Gusar Distr. (Gusar) [sub Trochosa c. Verzhbitsky, 1902], Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999]. COMMENTS. According to Platnick [2002] it has Palaearctic range, while we think it may have an Euro-Caucasian range.

Aulonia C.L. Koch, 1847 Aulonia albimana (Walckenaer, 1805)
Aulonia a.: Fuhn & Niculescu-Burlacu, 1971: 240, fig. 117ae, 118 (). A. a.: Roberts, 1985: 153, fig. 67a (). A. a.: Heimer & Nentwig, 1991: 324, fig. 857 (). A. a.: Roberts, 1995: 235, fig. (). A. a.: Roberts, 1998: 253, fig. ().

Arctosa leopardus (Sundevall, 1833)
Arctosa l.: Fuhn & Niculescu-Burlacu, 1971: 181, fig. 89ae (). A. l.: Roberts, 1985: 148, fig. 64c (). A. l.: Heimer & Nentwig, 1991: 320, fig. 854 (). A. l.: Roberts, 1995: 229, fig. (). A. l.: Roberts, 1998: 246, fig. (). Material examined: 35 5 juv. (YMT & CAS), SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001 (YM); 2 (YMT), SE Azerbaijan, ca 10 km W of Astara Town, Isti-Su, 38њ27N, 48њ47E, on the border with Iran, 25.04.2001 (YM).

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Dunin, 1989]. COMMENTS. This species seems to have an EuroCaucasian (?) range, while according to Platnick [2002] it has a Palaearctic distribution.

"Aulonia" kratochvili Dunin, Buchar et Absolon, 1986
Aulonia k. Dunin et al., 1986: 28, fig. 17 (). A. k.: Kronestedt, 1997: 77, fig. 12 (). Material examined: 24 juv. (YMT), SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 23 28.04.2001 (YM).

RECORDS: Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. According to Platnick [2002] it has Palaearctic range, while we think it may have an Euro-Central Asian range.

Arctosa perita (Latreille, 1799)
Artcosa p.: Holm, 1947: 20, fig. 6a, pl. 4, fig. 4041, pl. 9, fig. 25 (). A. p.: Lugetti & Tongiorgi, 1965: 175, fig. II.14, III, IV.1 (). A. p.: Fuhn & Niculescu-Burlacu, 1971: 185, fig. 91ad (). A. p.: Roberts, 1985: 148, fig. 64b (). A. p.: Heimer & Nentwig, 1991: 320, fig. 853 (). A. p.: Roberts, 1995: 228, fig. (). A. p.: Roberts, 1998: 246, fig. ().

RECORDS: Gobustan, Mingechaur, Yevlakh, Kyurdamir, Sheki, Saatly, Lenkoran [Dunin et al., 1986], Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. Known from Macedonia to Azerbaijan [Thaler et al., 2000]. While somatically this species is very similar to A. albimana (the genorotype), the male palp and female epigyne of this species differ considerably and clearly A. kratochvili belongs to another genus.

Evippa Simon, 1882 Evippa apsheronica sp.n.
See above. See above.

RECORDS: Apsheron Pen. [Dunin, 1984]. COMMENTS. According to Platnick [2002] it has a Holarctic range. In Eurasia it is known from western Europe to Caucasus [cf. Mikhailov, 1997].

Evippa caucasica sp.n.


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen Geolycosa Montgomery, 1904 Geolycosa dunini Zyuzin et Logunov, 2000
RECORDS: Nakhichevan Distr. (Nakhichevan) [sub Lycosa piochardi, Schmidt, 1895], Apsheron Pen. [sub L. narboniensis, Dunin, 1984], Sheki-Zagatala area [sub L. narbonensis, Dunin, 1989], Lenkoran area [sub L. nordmanni, Guseinov, 1999], Lerik Distr. (Gosmalyan), Yardymly Distr. (Avash), Salyan Distr. (Shirvan Reserve) [Zyuzin & Logunov, 2000]. COMMENTS. Distributed from Greece to Kyrgyzstan [Zyuzin & Logunov, 2000].

RECORDS: Khanlar Distr. (Gyanja) [sub Lycosa vultuosa, Schmidt, 1895], Sheki-Zagatala area [sub Lycosa vultuosa, Dunin, 1989], Lenkoran area [sub Lycosa vultuosa Guseinov, 1999], Shemakha Distr. (Pirkuli Reserve), Guba Distr. (Guba), Khanlar Distr. (Khanlar), Agdash (Agdash), Sheki Distr. (Sheki) [Zyuzin & Logunov 2000]. COMMENTS. This species has a Transcaucasian range [Zyunin & Logunov, 2000].

Geolycosa d. Zyuzin & Logunov, 2000: 309, fig. 79 ().

Mustelicosa Roewer, 1960 Mustelicosa dimidiata (Thorell, 1875)
RECORDS: Gusar Distr. (Gusar) [sub Trochosa d., Verzhbitsky, 1902]. COMMENTS. This species was never redescribed and illustrated, and therefore its record from Azerbaijan may refer to another species. M. dimidiata was described from environs of Volgograd and seems to be distributed from Volga to eastern China [cf. Marusik et al., 2000; Marusik & Logunov, 2002].

Hogna Simon, 1885 Hogna alticeps (Kroneberg, 1875)
RECORDS: (sub Lycosa alticeps): Apsheron Pen. [Dunin, 1984]. COMMENTS. It is known from Central Asia [Platnick, 2002]. As this species is known only from the female and has never been properly illustrated and it is probable that the records from Azerbaijan were based on misidentifications [cf. Mikhailov, 1997].
Tarentula a. Kroneberg, 1875: 40, pl. 4, fig. 28 ().

Pardosa C.L. Koch, 1847 Pardosa aenigmatica Tongiorgi, 1966
Pardosa a.: Tongiorgi, 1966a: 304, fig. 127 (). P. a.: Tongiorgi, 1968: 108, pl. II, fig. 13 ().

Hogna bergsoei (Thorell, 1875)
RECORDS: Apsheron Pen. [sub Lycosa b. L. Koch, 1878; sub Alopecosa b., Dunin, 1984]. COMMENTS. This species was transferred to Hogna by Roewer [1955]. Like most of the other wolf spiders described by Thorell it was never illustrated and redescribed and therefore its generic placement is uncertain. Because types of this species were never studied by L. Koch or Dunin, it is probable that records from Azerbaijan refer to other species.

RECORDS: Sheki-Zagatala area [Dunin, 1989]. COMMENTS. Known in Italy and Azerbaijan [Platnick, 2002], it is probable that the record from Azerbaijan refers to another species (e.g. Wadicosa sp.). We were unable to find specimens of this species in Dunins collection stored in ZMMU.

Pardosa agrestris (Westring, 1861)
Pardosa a.: Roberts, 1985: 134, fig. 55c, 57ab (). P. a.: Roberts, 1995: 214, fig. (). P. a.: Roberts, 1998: 228, fig. (). P. a.: Zyuzin & Logunov, 2000: 316, fig. 4344 ().

Hogna ocellata (L. Koch, 1878)
Lycosa o. L. Koch, 1878: 52, pl. 2, fig. 12 (). Hogna o.: Roewer, 1955: 249.

RECORDS: Apsheron Pen. [sub. Lycosa o., L. Koch, 1878; Arctosa o., Dunin, 1984]. COMMENTS. This species was transferred to Hogna by Roewer [1955]. It was never redescribed and properly illustrated and known so far only from the type locality, the Apsheron Pen. Location of its types remains uncertain.

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999]. COMMENTS. Palaearctic range [Platnick, 2002], while most probably it has an Euro-Altaian range. The record from Azerbaijan may refer to P. pontica [cf. Zyuzin & Logunov, 2000].

Hogna radiata (Latreille, 1817)
RECORDS: Khanlar Distr. (Gyanja) [sub Lycosa r., Schmidt, 1895], Sheki-Zagatala area [sub Lycosa r., Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999]. COMMENTS. Known from Mediterranean to Central Asia [Platnick, 2002]. The record from Central Africa [cf. Platnick, 2002] may refer to another species.
Lycosa r.: Fuhn & Niculescu-Burlacu, 1971: 195, fig. 95ae ().

Pardosa agricola (Thorell, 1856)
Pardosa a.: Roberts, 1985: 133, fig. 55b, 56 (). P. a.: Roberts, 1995: 213, fig. (). P. a. :Roberts, 1998: 227, fig. ().

RECORDS: Lenkoran area [Guseinov, 1999]. COMMENTS. Known from Europe to Kazakhstan.

Lycosa Latreille, 1804 Lycosa praegrandis C.L. Koch, 1836
Lycosa p.: Zyuzin & Logunov, 2000: 306, fig. 12 (). L. p.: Thaler et al., 2000: 1077, fig. 22, 2426, 2930, 40, 45 ().

Pardosa azerifalcataMarusik, Guseinov & Koponen, 2003
RECORDS: Ismailly Distr. [Marusik et al., 2003]. COMMENTS. This species is closely related and very similar to P. jerginiensis Ponomarev, 1979 and possibly was earlier misidentified.
Pardosa f. Marusik et al., 2003: in press.


Spiders (Arachnida, Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) Pardosa bifasciata (C. L. Koch, 1834)
Pardosa b.: Tongiorgi, 1966a: 292, fig. 811 (). P. b.: Fuhn & Niculescu-Burlacu, 1971: 81, fig. 33ad (). P. b.: Heimer & Nentwig, 1991: 328, fig. 878 (). P. b.: Roberts, 1995: 216, fig. (). P. b.: Roberts, 1998: 229, fig. ().

59

(YM); 1 3 (YMT), Azerbaijan, Ismailly Distr., Khanaya, 700 m, 0809.07.2001 (EG); 1 (YMT), Astara Distr., Istisu, 25.04.2001 (EG)

RECORDS: Shemakha Distr. (Pirkuli Reserve) [Guseinov, 2002]. COMMENTS. It seems that this species has an EuroMongolian range [cf. Marusik et al., 2000], and we think the record of this species from eastern China refers to P. hanrasanensis Jo & Paik, 1984 which was wrongly synonymised with P. bifasciata.

Pardosa buchari Ovtsharenko, 1979
Pardosa b. Ovtsharenko, 1979: 47, fig. 1415, 2831 (). P. b.: Buchar & Thaler, 1998: 709, fig. 1415 (). P. b.: Zyuzin & Logunov, 2000: 315, fig. 3133, 3739 (). Material examined: 1 (YMT), Ismailly Distr., Ismailly Reserve, 10001200 m, 14.07.2001 (EG)

RECORDS: Gusar Distr. (Gusar) [sub Lycosa annulata Verzhbitsky, 1902], Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. In Platnicks catalogue [2002] it is listed as a species with Palaearctic range. However judging from the figures (Fig. 26b in Saito, 1959) the record from Japan refers to another species. We think records of P. hortensis from south Siberia [cf. Mikhailov, 1997] are based on misidentifications. Study of numerous specimens of lycosids from Siberia and the Far East has not revealed this species and nor any other members of the proxima species group, so it seems that P. hortensis has an Euro-Caucasian range.

Pardosa incerta Nosek, 1905
Pardosa i.: Tongiorgi, 1966b: 339, fig. 1-2, 26 (, D). P. i.: Zyuzin & Ovtsharenko, 1979: 60, fig. 13, 67, 9, 11, 13 ().

RECORDS: Shemakha Distr. (Pirkuli Reserve), Dashkesan Distr. (Dashkesan) [Zyuzin & Logunov 2000]. COMMENTS. Until recently this species was not known in Azerbaijan [Mikhailov, 1997, 1998, 1999, 2000]. It was reported for the first time within Azerbaijan from the Pirkuli Reserve by Zyuzin & Logunov [2002]. P. buchari is so far known from northern Caucasus, Georgia and Azerbaijan.

RECORDS: Sheki-Zagatala area [Dunin, 1989]. COMMENTS. Known so far from Turkey, Azerbaijan and northern Caucasus (Kabardino-Balkaria, Krasnodar Prov.) only [Zyuzin & Ovtsharenko, 1979; Mikhailov, 1997].

Pardosa italica Tongiorgi, 1966
Pardosa i.: Tongiorgi, 1966a: 301, fig. 135138 (). P. i.: Fuhn & Niculescu-Burlacu, 1971: 95, fig. 40ae (). Material examined: 1 (YMT), Salyan Distr., Shirvan Reserve, 28.05.2000 (EG).

Pardosa caucasica Ovtsharenko, 1979
Pardosa c. Ovtsharenko, 1979: 48, fig. 1011, 2425 (). P. c.: Tцpfer-Hofmann, Cordes & von Helversen, 2000: 271, fig. 3233 (). Material examined: 1 (YMT), ca 70 km N of Baku, Galalty Vill., 40њ48N, 49њ12E, 19.04.2001 (YM).

RECORDS: Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. This species has a Caucasian range [Platnick, 2002].

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992]. COMMENTS. It seems that this species has an EuropeanCentral Asian range (cf. Mikhailov, 1997 & Platnick, 2002).

Pardosa jergeniensis Ponomarev, 1979
RECORDS: Apsheron Pen. (Shagan), Shemakha Distr. (Pirkuli Reserve) [Guseinov & Rubtsova, 2001]. COMMENTS. Until recently this species was known only from Kalmykia and northwestern Kazakhstan [cf. Mikhailov, 1997]. This species is very similar to P. azerifalcata Marusik et al., 2003 and possibly some or even all records of P. jergeniensis may refer to P. azerifalcata.
Pardosa j. Ponomarev, 1979: 1589, fig. 1 ().

Pardosa colchica Mcheidze, 1947
RECORDS: Khanlar Distr. (Gyoy-Gyol Reserve) [Zyuzin & Logunov 2000]. COMMENTS. Until recently this species was not known in Azerbaijan [Mikhailov, 1997, 1998, 1999, 2000]. It was reported for the first time within Azerbaijan from Khanlar Distr. by Zyuzin & Logunov [2002]. So far it is known from northern Caucasus, Georgia and Azerbaijan and has a transCaucasian range.
Pardosa c.: Zyuzin & Logunov, 2000: 310, fig. 1723 (, D).

Pardosa luctinosa Simon, 1876
Pardosa l.: Tongiorgi, 1964: 244, fig. 15 (). P. l.: Tongiorgi, 1966a: 300, fig. 139142 (). P. l.: Fuhn & Niculescu-Burlacu, 1971: 97, fig. 41ae ().

Pradosa gusarensis sp.n.
See above.

Pardosa hortensis (Thorell, 1872)
Pardosa h.: Tongiorgi, 1966a: 307, fig. 150151, 154155 (). P. h.: Roberts, 1985: 134, fig. 59c (). P. h.: Heimer & Nentwig, 1991: 332, fig. 1402 (). P. h.: Roberts, 1995: 220, fig. (). P. h.: Roberts, 1998: 234, fig. (). Material examined: 2 (YMT), SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001

RECORDS: Apsheron Pen. [Dunin, 1984], Muganskaya steppe [Dunin & Mamedov, 1992]. COMMENTS. Systematics of this species is obscure and it is unclear if it has a trans-Palaearctic range or is a complex species represented by series of subspecies or sibling species throughout Eurasia. Through the whole range this taxon is associated with salted lands.

Pardosa lugubris (Walckenaer, 1802)
Pardosa l.: Tцpfer-Hofmann & von Helversen, 1990: 349, fig. 1a ().


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen
l.: Roberts, 1998: 233, fig. (). l.: Kronestedt, 1999: 3, fig. 1c, 3fk, 5bc, 6b, 8b (). l.: Tцpfer-Hofmann, Cordes & von Helversen, 2000: 265, fig. ().

Pardosa paracolchica Zyuzin et Logunov, 2000
Pardosa p. Zyuzin & Logunov, 2000: 311, fig. 1316, 2427 (). Material examined: 1 , Azerbaijan, Ismailly Distr., Khanaya, 700 m, 08.07.2001 (EG).

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Atakishiev, 1969; Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. Study of material from Azerbaijan (by YM) reveals no true P. lugubris sensu Tцpfer-Hofmann et al. [2000] but P. caucasica and one possibly undescribed species. It is possible that records of P. lugubris are based on misidentifications and this species may not occur in Azerbaijan.

RECORDS: Guba Distr. (Nyugyadi) [Zyuzin & Logunov, 2000]. COMMENTS. Known only from Caucasus [Zyuzin & Logunov, 2000].

Pardosa pirculiensis Zyuzin et Logunov, 2000
Pardosa p. Zyuzin & Logunov, 2000: 314, fig. 2830, 3436 (). Material: Ismailly Distr., Ismailly Reserve, 1500 m, 12.07.2001 (EG).

Pardosa morosa (L. Koch, 1870)
Pardosa m.: Tongiorgi, 1966a: 311, fig. 6466 (). P. m.: Fuhn & Niculescu-Burlacu, 1971: 104, fig. 45ae (). P. m.: Buchar & Polenec, 1974: 83, fig. FI (). P. m.: Heimer & Nentwig, 1991: 332, fig. 1403 (). Material examined: 1 (ZMUT) Gabala Distr., Bum Vill., 05.06.2001 (EG).

RECORDS: Shemakha Distr. (Pirkuli Reserve) [Zyuzin & Logunov, 2000]. COMMENTS. Known only from Azerbaijan [Zyuzin & Logunov, 2000].

RECORDS: Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. Known from Europe to Central Asia [Platnick, 2002]. The single specimen that we have on hand fits well the illustration provided by Buchar & Polenec [1974] while differing from other illustrations. The color pattern of female from Bum Village corresponds well to those given by Buchar & Polenec [1974].

Pardosa pontica (Thorell, 1875)
Pardosa p.: Tongiorgi, 1966: 351, fig. 10-11, 24 (). P. p.: Fuhn & Niculescu-Burlacu, 1971: 118, fig. 53ac (). P. p.: Zyuzin & Logunov, 2000: 316, fig. 4042 ().

Pardosa nebulosa (Thorell, 1872)
Pardosa n.: Tongiorgi, 1966a: 303, fig. 119122 (). P. n.: Fuhn & Niculescu-Burlacu, 1971: 106, fig. 46ae (). P. n.: Heimer & Nentwig, 1991: 332, fig. 891 ().

RECORDS: Apsheron Pen. [Dunin, 1984], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999], Guba Distr. (Nyugyadi), Khanlar Distr. (Gey-Gyol Reserve), Apsheron Pen. (Baku), Lenkoran Distr. (Lenkoran), Shemakha Distr. (Pirkuli Reserve) [Zyuzin & Logunov, 2000]. COMMENTS. Known from Romania to Turkmenistan [cf. Mikhailov, 1997; Platnick, 2002].

RECORDS: Gusar Distr. (Gusar) [sub Lycosa n., Verzhbitsky, 1902], Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999]. COMMENTS. It has a Palaearctic range [Platnick, 2002]. Systematics and range of this species require investigation.

Pardosa proxima (C. L. Koch, 1847)
Pardosa p.: Tongiorgi, 1966a: 306, fig. 146147, 156157 (). P. p.: Fuhn & Niculescu-Burlacu, 1971: 122, fig. 55ae (). P. p.: Roberts, 1985: 134, fig. 59d (). P. p.: Heimer & Nentwig, 1991: 332, fig. 1405 (). P. p.: Roberts, 1995: 220, fig. (). P. p.: Roberts, 1998: 235, fig. (). Material examined: 1 1 , Gabala Distr., Laza, 12.08.2001 (EG) 1 , Azerbaijan, Ismailly Distr., Khanaya, 700 m, 09.07.2001 (EG).

Pardosa paludicola (Clerck, 1757)
Pardosa p.: Tongiorgi, 1966a: 295, fig. 2325 (). P. p.: Fuhn & Niculescu-Burlacu, 1971: 114, fig. 51ae (). P. p.: Roberts, 1985: 134, fig. 60b (). P. p.: Heimer & Nentwig, 1991: 332, fig. 890 (). P. p.: Roberts, 1995: 221, fig. (). P. p.: Roberts, 1998: 235, fig. ().

RECORDS: Lenkoran area [Guseinov, 1999]. COMMENTS. P. proxima is listed as a Palaearctic species [Platnick, 2002] but there are no proven records of this species in Siberia.

RECORDS: Sheki-Zagatala area [Dunin, 1989]. COMMENTS. Palaearctic range [Platnick, 2002].

Pardosa pullata (Clerck, 1757)
Pardosa p.: Holm & Kronestedt, 1970: 410, fig. 1, 4, 9, pl. I, fig. 12, pl. II, fig. 12; pl. III, fig. 1, pl. IV, fig. 12 (). P. p.: Roberts, 1985: 134, fig. 58b (). P. p.: Heimer & Nentwig, 1991: 330, fig. 885 (). P. p.: Roberts, 1995: 216, fig. (). P. p.: Roberts, 1998: 230, fig. ().

Pardosa palustris (Linnaeus, 1758)
Pardosa p.: Tongiorgi, 1966a: 283, fig. 8485, 91, 102103 (). P. p.: Fuhn & Niculescu-Burlacu, 1971: 116, fig. 52ae (). P. p.: Roberts, 1985: 134, fig. 57d, 58a (). P. p.: Heimer & Nentwig, 1991: 324, fig. 861 (). P. p.: Roberts, 1995: 215, fig. (). P. p.: Roberts, 1998: 229, fig. ().

RECORDS: Sheki-Zagatala area [Atakishiev, 1969]. COMMENTS. Trans-Palaearctic-Alaskan range [Marusik et al., 2000]. It is clear that the single record of P. lugubris for the whole Caucasus was based on a misidentification.

RECORDS: Sheki-Zagatala area [Atakishiev, 1969; Dunin, 1989]. COMMENTS. This species has an Euro-Baikalian range [cf. Mikhailov, 1997].

Pardosa saltuaria (L. Koch, 1870)
Pardosa s.: Tongiorgi, 1966a: 288, fig. 6771 ().


Spiders (Arachnida, Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae)
P. s.: Fuhn & Niculescu-Burlacu, 1971: 127, fig. 58ae (). P. s.: Wunderlich, 1984b: 430, fig. 2228 (). P. s.: Heimer & Nentwig, 1991: 328, fig. 873 ().

61

RECORDS: Lenkoran area [Guseinov, 1999]. COMMENTS. This species is known from western Europe to Kazakhstan [Platnick, 2002]. Records from Kazakhstan require confirmation.

RECORDS: Gusar Distr. (Gusar) [sub Lycosa v., Verzhbitsky, 1902], Sheki-Zagatala area [Dunin, 1989]. COMMENTS. Known from Europe to Caucasus [cf. Mikhailov, 1997 & Platnick, 2002].

P. v.: Thaler, 1987: 194, fig. 15 (). Material examined: 1 , Gabala Distr., Amirvan, 04.06.2001 (EG)

Pardosa schenkeli Lessert, 1904
Pardosa s.: Tongiorgi, 1966a: 291, fig. 1214 (T from Passiena). P. s.: Fuhn & Niculescu-Burlacu, 1971: 129, fig. 59ac (). P. s.: Heimer & Nentwig, 1991: 328, fig. 877 (). P. s.: Logunov & Marusik, 1995: 113, fig. 2024 ().

Pirata Sundevall, 1832 Pirata insularis Emerton, 1885
Pirata piccolo: Kronestedt, 1980b: 65, fig. 9I, K (). P. i. Dondale & Redner, 1990: 255, fig. 380386 (). P. piccolo: Heimer & Nentwig, 1991: 346, fig. 896 (). P. i.: Roberts, 1995: 234, fig. (). P. i.: Roberts, 1998: 251, fig. ().

RECORDS: Sheki-Zagatala area [Dunin, 1989]. COMMENTS. Trans-Palaearctic range [Marusik et al., 2000].

Pardosa sphagnicola (F. Dahl, 1908)
Pardosa s.: Holm & Kronestedt, 1970: 417, fig. 6, 11, pl. II, fig. 56, 13, pl IV, fig. 57 (). P. s.: Roberts, 1995: 218, fig. (). P. s.: Roberts, 1998: 231, fig. ().

RECORDS: Lenkoran area [Guseinov, 1999]. COMMENTS. Platnick [2002], treats P. insularis as having a Holarctic range while in fact in Eurasia it occurs only west of Yenisei, and has never been reported from Central, Eastern Siberia or Far East [cf. Mikhailov, 1997].

Pirata latitans (Blackwall, 1841)
Pirata l.: Roberts, 1985: 150, fig. 65e (). P. l.: Roberts, 1995: 234, fig. (). P. l.: Roberts, 1998: 252, fig. (). Material examined: 32 , SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001 (YM).

RECORDS: Gusar [Guseinov, 2002]. COMMENTS. This species has an Euro-Yenisei range [Marusik et al., 2002]. It seems that this species was misidentified with sibling P. pullata.

Pardosa tasevi Buchar, 1968
Pardosa t. Buchar, 1968: 122, fig. 4AB (). P. t.: Ovtsharenko, 1979: 49, fig. 1 (). P. t.: Weiss & Varvara, 1983: 273, fig. 1 ().

RECORDS: Sheki-Zagatala area [Dunin, 1989]. COMMENTS. This species so far is known only from Caucasus [Mikhailov, 1997].

RECORDS: Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. Known from Europe to Caucasus [Platnick, 2002].

Pirata piraticus (Clerck, 1757)
Pirata p.: Kronestedt, 1980b: 65, fig. 2A, 3B, D, 4A, 6AC, 7D F, 8EF, 9CD (). P. p.: Roberts, 1985: 150, fig. 65b, 66b (). P. p.: Dondale & Redner, 1990: 270, fig. 421424, 426427 (). P. p.: Heimer & Nentwig, 1991: 346, fig. 892 (). P. p.: Roberts, 1995: 232, fig. (). P. p.: Roberts, 1998: 250, fig. (). Material examined: 1 1 juv., SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 2328.04.2001 (YM)

Pardosa tatarica (Thorell, 1875) Figs. 4750.
Pardosa strigillata: Tongiorgi, 1966: 308, fig. 143144, 158 159 (). P. strigillata: Zyuzin, 1979: 435, fig. 12, 49 (). P. strigillata: Heimer & Nentwig, 1991: 332, fig. 1401 (). Material examined: 1 , Gabala Distr., Bum Vill., 05.06.2001 (EG); 9 4 , SE Azerbaijan, ca 10 km W of Astara Town, IstiSu, 38њ27N, 48њ47E, on the border with Iran, 25.04.2001 (YM). Comparative material: 2 2 (YMT), GREECE, Trace, Samothrace Isl., Therma, stream bank, 13.04.1996 (leg. P. Lindskog, det. T. Kronestedt).

RECORDS: Sheki-Zagatala area [Dunin, 1989], Apsheron Pen. (Bina) [Guseinov & Rubtsova, 2001]. COMMENTS. Holarctic range [Platnick, 2002].

RECORDS: (sub P. strigillata): Azerbaijan [no precise locality, Zyuzin, 1979], Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. Known from Europe to Caucasus [cf. Mikhailov, 1997 & Platnick, 2002]. Comparison of populations from Greece and Azerbaijan (cf. Figs. 4750) suggests that specimens from Azerbaijan belong to another species. Specific placement of the Azerbaijanian population will be solved when we study types or topotypes of P. tatarica that might be not conspecific with P. strigillata.

Trochosa C.L. Koch, 1847 Trochosa hispanica Simon, 1870*
Trochosa h.: Hдnggi, 1989: 168, fig. 1ab (). T. h.: Thaler et al., 2000: 1079, fig. 3739, 41. Material examined: 4 3 2 juv., SE Azerbaijan, Lenkoran Distr., env. of Aurora Vill., 38њ40N, 48њ52E, 23 28.04.2001 (YM); 2 1 juv., SE Azerbaijan, ca 10 km W of Astara Town, Isti-Su, 38њ27N, 48њ47E, on the border with Iran, 25.04.2001 (YM)

Pardosa vittata (Keyserling, 1863)
Pardosa v.: Tongiorgi, 1966a: 292, fig. 17 (). P. v.: Fuhn & Niculescu-Burlacu, 1971: 134, fig. 62ae ().

COMMENTS. This species can be easily recognized in the field by the stripe of white hairs on the dorsal side of the males tibia I [Thaler et al., 2000], in alcohol these hairs sometimes can be poorly visible.


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Yuri M. Marusik, Elchin F. Guseinov & Seppo Koponen

Figs. 5156. Copulatory organs of Pardosa sp. (51) and Wadicosa fidelis (O. Pickard-Cambridge, 1872)? (5256). 51, 56 epigyne, ventral view; 5254 palp, dorsal, pro- and retrolateral view respectively; 55 tip of cymbium with two claws. Scale = 0.1 mm except for Fig. 55 (=0.05 mm). Figs. 5156. Копулятивные органы Pardosa sp. (51) и Wadicosa fidelis (O. Pickard-Cambridge, 1872)? (5256). 51, 56 эпигина, вид снизу; 5254 пальпа самца, разные аспекты; 55 вершина цимбиума с двумя коготками. Масштаб 0,1 мм за исключением рис. 55 (0.05 мм).

Previously this species was reported from Spain to Russia [Platnick, 2002], while in fact its correct range is Spain to Ukraine, or Spain to Tajikistan. T. ruricola rustica Thorell, 1875 a junior synonym of T. hispanica was described from Italy: Gennazano () and Russia (=Russian Empire): Yekaterinoslav (=Dnipropetrovsk, Ukraine now) () and since then was reported only from Tajikistan by Charitonov [1951]. This species is new for Azerbaijan and Caucasus as a whole.

COMMENTS. This species has a trans-Palaearctic range [Platnick, 2002]. Records of this species, at least those from Lenkoran, may refer to T. hispanica.

Trochosa spinipalpis (F.O. Pickard-Cambridge, 1895)
Trochosa s.: Fuhn & e (). T. s.: Roberts, 1985: T. s.: Roberts, 1995: T. s.: Roberts, 1998: Niculescu-Burlacu, 1971: 229, fig. 112a 144, fig. 62f, 63d (). 228, fig. (). 245, fig. ().

Trochosa ruricola (De Geer, 1778)
Trochosa r.: Fuhn & f (). T. r.: Roberts, 1985: T. r.: Roberts, 1995: T. r.: Roberts, 1998: Niculescu-Burlacu, 1971: 226, fig. 111a 144, fig. 62c, 63a (). 226, fig. (). 244, fig. ().

RECORDS: Khanlar Distr. (Gyanja) [sub Lycosa r., Schmidt, 1895], Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999].

RECORDS: Apsheron Pen. [Dunin, 1984], Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992], Lenkoran area [Guseinov, 1999]. COMMENTS. This species has a trans-Palaearctic range [Platnick, 2002]. Records of this species, at least those from Lenkoran, may refer to T. hispanica.


Spiders (Arachnida, Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) Trochosa terricola Thorell, 1856
Trochosa t.: Fuhn & Niculescu-Burlacu, 1971: 231, fig. 113ae (). T. t.: Roberts, 1985: 144, fig. 62e, 63c (). T. t.: Roberts, 1995: 227, fig. (). T. t.: Roberts, 1998: 245, fig. ().

63

RECORDS: Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. This species has a circum-Holarctic range [Platnick, 2002]. Records of this species, at least those from Lenkoran, may refer to T. hispanica.

Wadicosa Zyuzin, 1985
This genus was only recently reported from Azerbaijan and Caucasus as a whole [Marusik & Guseinov, 2003].

longing to 14 genera: Allohogna (1 species), Alopecosa (14, of which 4 species doubtful), Arctosa (4), Aulonia (2), Evippa (2), Geolycosa (1), Hogna (4, 2 species doubtful), Lycosa (1), Mustelicosa (1 species doubtful), Pardosa (28, 4 species doubtful), Pirata (3), Trochosa (4), Wadicosa (1), Xerolycosa (2). Of them, one genus and 11 species records are doubtful and require confirmation. Taking into consideration that many areas of Azerbaijan such as highlands in Gusar and Zagatala Districts, mountains of Talysh area, Nakhitchevan are very poorly studied, and pitfall trapping was rarely used, it is reasonable to expect many new species and even few generic discoveries.
ACKNOWLEDGEMENTS. We wish to thank Galina Azarkina (Novosibirsk), Torbjцrn Kronestedt (Stockholm), Dmitry Logunov (Manchester) who gave us comparative material or types deposited in their museums. We are highly indebted to T. Kronestedt, Vladimir I. Ovtsharenko (New York) and Pekka T. Lehtinen (Turku), who supplied us with important literature. Special thanks are extended to Izyaslav M. Kerzhner (St. Petersburg) for his consultation on nomenclatorial problems. We are also grateful to D.J. Buckle (Saskatoon) for checking English. This work was supported in part by the Russian Foundation for Basic Research (grant # 01-0448989) and the Academy of Finland (grant # 79224).

Wadicosa fidelis (O. Pickard-Cambridge, 1872) ? Figs. 5256.
Wadicosa f.: Song et al., 1999: 202C, G (). Material examined: 3 2 2 juv. (YMT), south-eastern Azerbaijan, ca 10 km W of Astara Town, Istisu, 38њ27N, 48њ47E, on the border with Iran, 25.04.2001 (YM).

COMMENTS. This species has the widest distribution among the genus and is known from western Europe to Japan. Our identification was confirmed with some uncertainty by T. Kronestedt. Figures provided to this species by different authors [e.g. Wunderlich, 1992: fig. 727728 and Song et al., 1999] seem to belong to two different species, so W. fidelis may have a smaller range and be restricted to the western Palaearctic. Our specimens fit well the figures provided by Song et al. [1999].

References
Alderweireldt M. 1991. A revision of the African representatives of the wolf spider genus Evippa Simon, 1882 (Araneae, Lycosidae) with notes on allied species and genera // J. Nat. Hist. Vol.25. P.359381. Alderweireldt M. 2002. Redescription of Arctosa tbilisiensis Mcheidze, 1947, a poorly known Pontocaspian wolf spider (Araneae, Lycosidae) // Bull. Br. arachnol. Soc. Vol.12. No.4. P.183184. Atakishiev T.A. 1969. [Spiders symbionts of the honey bee] // Uchenyye zapiski Kazanskogo veterinarnogo instituta. Vol.105. P.317323 [in Russian] Bonnet P. 1955. Bibliographia Araneorum. Vol.2. Part 1. Toulouse. P.1918. Buchar J. 1968. Zur Lycosidenfauna Bulgariens (Arachn., Araneae) // Vestn. Иs. spoleи. zool. Bd.32. S.116130. Buchar J., Polenec A. 1974. Zur Lycosidenfauna Jugoslawiens (Araneae: Lycosidae) // Vestn. Иs. spoleи. zool. Bd.38. S.81 85. Buchar J., Thaler K. 1998. Lycosidae from the high alpine zone of the Caucasus range, with comparative remarks on the fauna of the Alps (Arachnida: Araneae) // Linzer biol. Beitr. Bd.30. P.705717. Charitonov D.E. 1932. Katalog der russischen Spinnen. Leningrad: AN SSSR Publ. 206 S. Charitonov D.E. 1951. [Spiders and harvestmen of the Kondara Gorge] // Ushchelye Kondara. Moscow-Leningrad. P.209 216 [in Russian]. Dondale C.D., Redner, J.H. 1990. The wolf spiders, nurseryweb spiders, and lynx spiders of Canada and Alaska (Araneae: Lycosidae, Pisauridae, and Oxyopidae) // The Insects and Arachnids of Canada. No.17. 383 pp. Dunin P.M. 1984. [Fauna and ecology of the spiders of the Apsheron Peninsula, Azerbaijan SSR] // Fauna i ekologiya paukoobraznykh. Perm. P.4560 [in Russian]. Dunin P.M. 1989. [Fauna and altitudinal distribution of spiders (Arachnida, Aranei) of the Azerbaijan part of the southern macroslope of the Caucasus Major] // Fauna i ecol. paukov i skorpionov. Moscow: Nauka. P.3139 [in Russian].

Xerolycosa F. Dahl, 1908 Xerolycosa miniata (C.L. Koch, 1834)
Xerolycosa m.: e (). X. m.: Roberts, X. m.: Roberts, X. m.: Roberts, Fuhn & Niculescu-Burlacu, 1971: 235, fig. 115a 1985: 142, fig. 61b (). 1995: 223, fig. (). 1998: 237, fig. ().

RECORDS: Sheki-Zagatala area [Dunin, 1989], Muganskaya steppe [Dunin & Mamedov, 1992]. COMMENTS. This species has an Euro-Yenisei range [Marusik et al., 2000].

Xerolycosa nemoralis (Westring, 1861)
Xerolycosa n.: Fuhn & Niculescu-Burlacu, 1971: 237, fig. 116a e (). X. n.: Zyuzin, 1985a: 48, fig. 15-16, 2022 (). X. n.: Roberts, 1985: 140, fig. 61a (). X. n.: Roberts, 1995: 222, fig. (). X. n.: Roberts, 1998: 236, fig. ().

RECORDS: Sheki-Zagatala area [Dunin, 1989], Lenkoran area [Guseinov, 1999]. COMMENTS. This species has a trans-Plaearctic range [Marusik et al., 2000].

Conclusions
Taking into account the new data given above, the lycosid fauna of Azerbaijan encompass 68 species be-


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