Документ взят из кэша поисковой машины. Адрес оригинального документа : http://www.mce.biophys.msu.ru/archive/doc15826/doc.pdf Дата изменения: Tue Oct 30 16:24:56 2007 Дата индексирования: Mon Oct 1 20:51:32 2012 Кодировка:

.. (. , ) ­ ­ , . , . MODEL OF THE RHYTHMICAL SPOROGENESIS IN STREPTOMYCES COLONIES Saveliev A. P. (Puschino, Moscow Region) It is considered single measured imitation model of the growth and zoneforming of the radiant fungi ­ streptomycetes ­ colony on synthetic agar nutritious with piruvate, glucose or succinate as a limiting source of energy and carbon. A certain basic and conducting parameters was determine. The quantities of this parameters is set by composition of nutritious and condition of environmental space. [1-5, 7-12], , , .. , : · , , () ,
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; · , ; · ( ) ( , ) - ; · ( ) ( .. ) , . , . Streptomyces levoris Kras., : , 2 40 mmol/l [1-4, 14]. "" +27 C. 2-3 . [6]. , 10-15% [14]
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[1-5, 7-12] : , , . . . , , . , , : ) ; ) ) [1]. , , , . , ­ , , , . , ) , ; ) ; ) - - - , ; ­ . .
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0.5 10-15 ( ), , . [5,6,10], (.1) : R=Kr*t (1) R ­ (), Kr ­ (/), t ­ , (). (.1), [12,13]: R=Kr*(t ­ ()) (2) () ­ 2-, 3- - . , ( ) n- - : R n,p = R1 + *(n­1) ­ *(­1) (3) t n,p = t1 + T*(n­1) + *(­1) (4) R1 ­ , ­ , n ­ , ­ , ­ , t1 ­ , T ­ , ­ , . : Kr = ro / o (5) ro ­ , o ­ , .
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Kr [S] , [10]: Kr=Kr*[(S+So)/S]*{[(Skr+So)2Ks+So+So2Ki]/[(Skr+So)2Ks+(S+So)+(S+So)2 Ki]} (6) Kr- S=0; Skr ­ -, Kr (mmol/l), So ­ -, Kr=0, Ki , Ks- . - 1:
Kro 0.3 0.3 0.3 Sk 1.5414 16.256 1.0562 So 0.5336 3.958 0.0686 Ks 0.1244 2.5 0.0689 Ki 0.0134 5.798 0.0119 0.272 0.284 0.216

ro = *Kr (7) ­ , ro=f([S]) Kr: ­ , , [15]. , , ­ . . [8] , : Wn = (t ­ tn)*W^/(w+(t ­ tn)) (8) W^ ­ () (), (t ­ tn) ­ (), w ­ ­ (), W^. (8) , w ­ "" , . , , , 227


2. ,

, . (1), (3), (8) : R = t*Kr Rn,p = ro**(n -(-1)/P) tn,p = o*(*n+ +1+v*(-1)) Wn = = tz*W^/(w+tz) R , = .() = 0.5* Wn ± Rn,p . ()
. 1. : R ­ (); Kr ­ (/); t ­ ( ) (); Rn,p ­ n- (); tn,p ­ n- (); n = 1,2,3,...,N ­ , ; ­ ; = 1,2,3,...,P ­ ; ro ­ (); o ­ ; v ­ ; ­ ; Wn ­ (); W^ ­ () (); tz = (t ­ tn) ­ n- (); w ­ "" ; R, ­ () () n- ().

(4) /9/ /10/ /11/ /12/ /13/

, Kr, ro, , v .. , [1-4, 14-17]. v , , ()
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. : , , .

. 2.

. 2 , 5 mmol/l . (% ­ ) . . 2:
[S] t (mmol/l) () 5 12 5 12 5 12 N 5 6 4 P r () Kr v W^ w (/) () () () 2 0.144 0.5069 8 4 1.2 5 1.25 3 0.1397 0.5136 6 2 2.0 1 0.3 3 0.1365 0.6323 11 3 0.65 1 0.6

, . , "" , , , , , [8]. , .. ,
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­ , , , . , . , , : · ; · , , ; · , , , , , . , , , , , ; · - , : , Kr, ro . . · . 1. .X. ., . . .1976..31-37. 2. .. .
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3. 4. 5. 6. 7. 8. 9. 10. 11. 12. 13. 14. 15. 16. 17.

. , . 1976. .38-51. .., .., . . . 1978. .21-25. .., .., . . . 1978. .26-35. .., .., . 1982. .51. .6. . 954-960. .., . . «».1978.331. ., ., . . "".1980. Saveliev A. P., Studia biophysica. 1984. V.102. 1. P.51-60. Saveliev A. P., Studia biophysica. 1984. V.102. 1. P.61-67. Saveliev A. P., Gen. Phys. and Biophys. 1987. V.6. P.79-85. .., .., . 1999. .44. .2. .318-324. .., .., .., .., .., II . . . 1999. .278-279. .., .., .., .., .., . 1999. .44. .3. .505-509. .., .X., .., 1981. .6. .28-30. .., . . .: 1983. 131. .., , . . 1996. 4. . 460-466. .., .., II . . . 1999. .911-912.

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