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Arthropoda Selecta 16 (1): 1517

ї ARTHROPODA SELECTA, 2007

A new species of the mygalomorph spider genus Brachythele Ausserer, 1871 (Aranei: Nemesiidae) from Greece Новый вид мигаломорфных пауков рода Brachythele Ausserer, 1871 (Aranei: Nemesiidae) из Греции S.L. Zonstein С.Л. Зонштейн
Department of Zoology, The George S. Wise Faculty of Life Sciences, Tel-Aviv University, 69978 Tel-Aviv Israel. Email: znn@post.tau.ac.il Отдел зоологии, Факультет естественных наук им. Дж. С. Визе, Тель-Авивский университет, 69978 Тель-Авив Израиль.

KEY WORDS: Aranei, spiders, Nemesiidae, Brachythele, Greece.
КЛЮЧЕВЫЕ СЛОВА: Aranei, пауки, Nemesiidae, Brachythele, Греция.

ABSTRACT. B. bentzieni sp.n., can be distinguished from related species by the unique configuration of a shortened and keeled embolus, is described from a single male. РЕЗЮМЕ. Приведено описание самца B. bentzieni sp.n., отличающегося от близких видов уникальной конфигурацией укороченного и снабженного килем эмболюса самца. The Mediterranean representatives of Brachythele Ausserer, 1871 include 7 species, distributed from North- and North-Western Balkans to Cyprus and South-Western Turkey (see Platnick, 2007). At least one more species, whose description is given below, occurs in Eastern Greece. The following abbreviations applied in spider systematics are used: ALE anterior lateral eyes, AME anterior median eyes, PLE posterior lateral eyes, PME median lateral eyes; PLS posterior lateral spinnerets, PMS posterior median spinnerets; Fm femur, Pt patella, Tb tibia, Mt metatarsus, Tr tarsus; M megaspine; d dorsal, v ventral, p prolateral, r retrolateral positions; pd, rd, pv, rv the corresponding intermediate states. The number of some symmetrically located structures such as cheliceral teeth and denticles, and maxillary cuspules is shown as the number of structures located left of the longitudinal body axis (from an observers point of view) plus a corresponding number on the right side, respectively. The total body length measured does not include the length of the chelicerae. All measurements are given in millimeters, except eye diameters/interspaces shown as a ratio of the microscope scale units. All references are given according to Platnick [2007].

Brachythele bentzieni sp.n. Figs 18.
TYPES. holotype labeled: 14038, Brachythele icterica C. K., Volo (St. Nll.), Brachythele new species, det. M. Bentzien, Dec. 1975; dep. MNHN.

DIAGNOSIS. The new species can be easily distinguished from males of B. icterica (C.L. Koch, 1838), B. media Kulczyсski, 1897, B. denieri (Simon, 1916) and B. varrialei (Dalmas, 1921), which share a uniformly tapering embolus, by the presence of a well developed embolic keel (sf. Koch, 1838, Abb. 351; Drenski, 1937, fig. 8; Poleneи, 1978, fig. 2; Raven, 1985, fig. 89). The same character is found in a recently described Bulgarian congener, B. langourovi Lazarov, 2005, but it differs from this species by the noticeably shorter and less curved embolic portion of the bulb and by the shape of the male embolic keel. It differs from two other congeners with unknown males as follows: from B. incerta Ausserer, 1871 by the strongly recurved fovea (that is almost straight in the former species) as well as by the presence of a distinct cheliceral rastellum, and from B. specularix Kulczynski, 1897 by fewer cheliceral teeth (7, as in the most members of the genus, vs. 89) and by the larger number of teeth on the paired tarsal claws (57 vs. 34). MALE (holotype). Total body length ca. 9.00. Carapace: length 4.90, width 3.93. Color in alcohol: carapace, palps and legs dorsally brown-rufous; eye tubercle brown; chelicerae reddish-brown; sternum, labium, maxillae, palps and legs ventrally light brown; abdomen dorsally light brown with a darker brown dorsal pattern represented by five pairs of inclined chevron-like fasciae, ventral abdominal surface and spinnerets pale yellowish brown. Carapace covered with moderately dense short and fine semi-appressed discolored hairs. Ratio of AME, ALE, PLE, PME: 7, 11, 9, 7. Interspaces: AMEAME 5, ALEAME 4, ALEPLE 3, PLE PME 3, PMEPME 14. Chelicerae with 7+7 promarginal teeth and 10+9 denticles, weak rastellum consists of 2530 stout setae. Labium: length 0.33, width 0.95; cuspules absent. Maxillae: 24+19 cuspules, serrula absent. Sternum: length 2.57, width 1.97. Sternal sigillae small rounded submarginal. Palp: Fm d1.1.1, pd1, Pt 0; Tb d1.1.1, p1.1, v1.1.1.


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S.L. Zonstein

Figs 18. Brachythele bentzieni sp.n., male: 1 carapace dorsally; 2 serrate cheliceral fang retrolaterally; 3 eye tubercle dorsally; 4 sternum, labium and maxillae ventrally; 5 right tibia and metatarsus I retrolaterally; 6 right tibia I ventrally; 7 left palpal tibia, cymbium and bulb ventrally; 8 same structures retrolaterally. Scale bars for Figs 1, 2 and 4 1.0 mm, and for Figs 3, 58 0.5 mm. Orig. Рис. 18. Brachythele bentzieni sp.n., самец: 1 карапакс сверху; 2 пильчатый коготок хелицеры ретролатерально; 3 глазной бугорок сверху; 4 стернум, лабиум и максиллы снизу; 5 правые голень и предлапка I ретролатерально; 6 правая голень I снизу; 7 левые голень пальпы, цимбиум и бульбус снизу; 8 те же структуры ретролатерально. Масштаб: 1, 2 и 4 1,0 мм; 3, 58 0,5 мм. Ориг.


A new species of Brachythele (Aranei: Nemesiidae) from Greece
Leg I: Fm d1.1.1.1, pd1.1; rd1.1. Pt p1.1; Tb p1.1, r1, v1.2.1+2M; Mt p1.1, v1.2.2. Leg II: Fm d1.1.1.1.2; pd1.1, r1.1.1; Pt p1.1; Tb p1.1, v2.2.3; Mt p1.1.1; v1.2.2. Leg III: Fm d1.1.1.1.2, pd1.1.1, rd1.1.1; Pt p1.1.1, r1.1; Tb d1, p1.1, r1.1.1, v2.2.2; Mt d1.1.1, p1.1.1, r1.1.1, v2.2.3. Leg IV: Fm d1.1.1.1, pd1.1.1, pr1.1.1; Pt 0; Tb d1, r1.1.1.1, v2.2.2; Mt d2.2.2, p1.1.1.2.1, r1.1.1, v2.2.2. Leg tarsi and cymbium aspinose. Leg measurements as shown in Table. Scopula: distal on metatarsi III, entire on tarsi III, vestigial on tarsi III, absent on tarsi IV. Paired claws: outer margin with 56 teeth, 7 teeth on inner margin. Trichobothria: 2 rows of 911 on tibiae, 1415 on metatarsi, 1314 on tarsi, 9 on cymbium. PMS: length 0.37, diameter 0.27. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.60, 0.47, 0.50; total length 1.57; apical segment triangle. Leg measurements:

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ACKNOWLEDGEMENTS. I am greatly obliged to the Late Dr. Jacqueline Heurtault, the former curator of arachnid collection at Musйum national dNistoire naturelle, Paris (MNHN), for allowing me to examine the holotype in 1984. I use this opportunity to thank Dr. Christo Deltchev for providing some comparative material from Bulgaria for examination. The study was completed with financial help provided generously by staff members of the Ministry of Absorption, Israel.

References
Drensky P. 1937. Faounata na paiatzite (Araneae) w Beulgaria. Podrazred Mygalomorphae: semeystwa Ctenizidae i Atypidae // Izw. tzar. prirod. Inst. Sofia. Vol.10. P.259280. Koch C.L. 1838. Die Arachniden. Nьrnberg. Bd.5. S.1104. Lazarov S. 2005. A new species from Bulgaria, Brachythele langourovi sp. n. (Araneae, Nemesiidae) // Rev. Suisse Zool. Vol.112. No.1. P.189193. Platnick N.I. 2007. The world spider catalog, version 8.0. American Museum of Natural History, online at http://research.amnh. org/entomology/spiders/catalog/index.html Poleneи A. 1978. Zusammensetzung und Besonderheiten der Epigaischen Spinnenfauna des Seslerio-Ostrietum am Berge Slavnik (1028 m) (Nord-Istrien, Jugoslavien) // Symp. Zool. Soc. London. No.42. P.367377. Raven R.J. 1985. The spider infraorder Mygalomorphae (Araneae): cladistics and systematics // Bull. Amer. Mus. Nat. Hist. Vol.182. No.1. P.1180.

Femur 2.14 3.80 3.50 3.16 3.78 Patella 1.02 2.10 1.91 1.56 2.04 Tibia 1.55 2.46 2.15 1.98 2.39 Metatarsus 2.78 2.44 2.91 1.84 Tarsus 0.8 2 2.20 1.7 8 3.94 2.0 0 Total 5.53 13.35 11.78 11.45 14.15 FEMALE. Unknown. DISTRIBUTION. Known presently only from Eastern Greece: Volos. ETYMOLOGY. Named after Mr. Michael Bentzien, an American MS-student, who first examined the specimen and referred it to a undescribed species.