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Number 65: 1-9

ISSN 1026-051X

November 1998

NEW DATA ON THE GENUS BRACHYOPA MEIGEN, 1822 (DIPTERA, SYRPHIDAE) FROM RUSSIAN FAR EAST V. A. Mutin
Department of Zoology, Komsomolsk-na-Amure State Pedagogical Institute, Komsomolsk-na-Amure, 681000, Russia Review of 12 species of the genus Brachyopa from Russian Far East is given. Brachyopa primorica sp. n. and B. zhelochovtsevi sp. n. are descibed. KEY WORDS. Syrphidae, hover-flies, taxonomy, distribution, new species. .. . Brachyopa Meigen, 1822 (Diptera, Syrphidae) // . 1998. N 65. C. 1-5. 12 Brachyopa . : Brachyopa primorica sp. n. B. zhelochovtsevi sp. n. -- , , --, 681000, . INTRODUCTION The genus Brachyopa (sensu Stackelberg, 1970) is distributed in Holarctic region. There are three centers of species diversity: European, Nearctic and Russian 1


Far East. The known larvae of this genus either occur in sap runs or in the accumulations of decaying sap under bark of leaf-bearing trees mostly. Twelve species of the genus Brachyopa including two new ones are reviewed from Russian Far East below. LIST OF THE SPECIES Brachyopa primorica Mutin, sp. n. Figs 1-4 MATERIAL. Holotype - , Primorskii krai: 30 km N Ternei, 3.VI 1982 (V. Mutin), deposited in the Institurte of Biology and Pedology (Vladivostok). Paratypes: 1, with the same data as holotype; Primorskii krai: 3, 4, Kamenushka, 30.V - 4.VI 1989 (A. Shatalkin); Khabarovskii krai: 3, Komsomolsk-na-Amure, 9.VI 1996 (D. Gritskevich); 2, 1, the same locality, 8.VI 1998 (V. Mutin); 1, the same locality, from larvae on bark of Ulmus pumila, IX 1996 (V. Mutin); 1, the same locality, IX 1997 (D. Gritskevich); 5, the same locality, 30.IX. 1997 (A. Sivova). DESCRIPTION. Male. Length: body - 7-8 mm, wing - 7-8 mm. Face yellow, pollinose, with bare more or less large shining spots under antennae. Gena shining yellow or brownish, with white pile. Frons yellow, bare, pollinose along eyes, with medial depression. Vertex pollinose in anterior half and black around ocelli, with short pale pile. Occiput mostly grey pollinose, with white pile. Antennae orangeyellow; flagellomere with very small and indistinct sensory pit; arista brownish, bare. Scutum dark grey, with pale grey pollinose vittae and maculae, submedial vittae are widest and connected posteriorly with large macula before scutellum. Scutum pile subpressed short mainly black, except pale pile of postpronotum and anterior part of scutum. Postalar callus brownish, with black setae. Scutellum yellow or brownish, with short subpressed mainly black pile and a few long black setae on posterior margin. Pleuron pale pilose, dull grey, except katatergum (pleurotergite) yellow or brownish; posterior anepisternum (hind part of mesopleuron) with a few long posterodorsal black setae. Wing mostly clean, with dark apex; pterostigma yellowish. Legs brownish yellow, two apical tarsomeres black. Abdomen shining black, subpressed white pilose; tergum 1 partly dull brownish; tergum 2 pollinose basally. Sterna 1-4 dull brownish-grey. Sterna 6-8 shining reddish-yellow, pale pilose. Genitalia - Figs 1-4. Female. Length: body - 7.8-8 mm, wing - 7,8 mm. Similar to male except of sexual characters. Face and gena shining yellow or reddish, narrow pollinose along eyes, with short pollinose fascia under antennae. Frons shining yellow or reddish, with triangular pillose maculae in posterior part. Vertex rather dull reddish, 2


Figs 1-5. Male of Brachyopa. 1-4) B. primorica sp. n.: 1) theca of hypandrium, lateral aspect, 2) theca of hypandrium, ventral aspect, 3) aedeagus, lateral aspect, 4) surstylus and cercus, lateral aspect, 5) B. cinerea, theca of hypandrium, lateral aspect. with short subpressed yellow pile. Apical half of tergum 2 and terga 3-5 entirely shinning dark brown. DIAGNOSIS. The new species resembles B. cinerea but differs by dark apex of the wing, by yellow katatergum, by shining outside part of postpronotum as well as by totally yellow frons of female and especially by shape of male hypandrium (Fig. 1 vs. Fig. 5). DISTRIBUTION. Russia: southern part of Khabarovskii krai, Primorskii krai. 3


Brachyopa zhelochovtsevi Mutin, sp. n. Figs 6-9 MATERIAL. Holotype - , Magadanskaya oblast': Ust'-Omchug, 23.VI 1963 (Zhelochovtsev), deposited in the Zoological Museum of Moscow State University. Paratypes: 3, the same locality, 24-30.VI 1963 (Zhelochovtsev); 2, Khabarovskii krai: Myaochan Range, Gornyi, 8.VI 1988, 6.VII 1991 (V. Mutin); 1, Myaochan Range, Chalba River, 2.VII 1979 (V. Mutin); 3, 20 km N Komsomolsk-naAmure, 10,13.VI 1983 (V. Mutin); 1, NW Amurskaya oblast': Nyukzha River, 3.VII 1976 (R. Soboleva). DESCRIPTION. Male. Length: body - 6-7.5 mm, wing - 6-7 mm. Face yellow, pale pollinose, with narrow shining medial stripe in lower part. Gena shining yellow, usually restricted by darken stripe anteriorly. Frons bare, rather shining yellow, with indistinct furrows. Vertex pale pollinose anteriorly, brownish or reddish around ocelli. Occiput dense pale pollinose and pale pilose. Antenna orange yellow; flagellomere with large ventrolateral roundish sensory pit; its diametr usually larger than distance between its margin and lower margin of antenna; arista long pilose, brown, yellow basally (Fig 6). Thorax reddish-yellow or brownish. Scutum mainly dark brown pilose, with a touch of pale pile in anterior part and black rather short setae laterally, with wide submedial vittae of pale polline, indistinct lateral ones and very narrow medial one, which curved to large macula before scutellum. Postpronotum paler, more or less shinning outside. Pleuron mostly pollinose, rare pale pilose, with 5-7 black setae on posterodorsal part of posterior anepisternum (posterior part of mesopleuron). Katepisternum (sternopleuron) and meron (hypopleuron) sometimes partly dark brown. Scutellum large, trapezoid, with short subpressed coarse black pile and apical rather short black setae. Wing clean. Femora yellow or brownish; tibia slightly darker than femora; tarsus brown dorsally, with darker apical tarsomeres. Terga 2,3 and 4 yellow, mainly short pale pilose, usually with dark medial vitta and posterior margin of terga. Sterna 7 and 8 yellow, pale pilose. Genitalia - Figs 7-9. Female. Length: body - 6.5-7.5 mm, wing - 6.5-8 mm. Similar to male except sexual characters. Face pollinose only under antennae, shining in lower projected part. Frons over antennae bare, shining yellow; posterior part of frons and vertex dense pollinose, with short pale pile. Sensory pit of flagellomere very large, reniform or triangular (Fig 6). Abdomen sometimes totally yellow or with darkish posterior margin of terga only. DISTRIBUTION. Russia: Magadanskaya oblast', Khabarovskii krai, Amurskaya oblast'. DIAGNOSIS. The new species is similar to B. testacea, but differs by larger trapezoid scutellum, by large sensory pit (Fig. 6 vs. Fig. 10) and by form of hypandrium tecka (Figs 8, 9 vs. Figs 11, 12). The same characters differ new species from other similar species, B. obscura Thompson et Tore, 1982. 4


Figs 6-13. Hover-flies. 6-9) Brachyopa zhelochovtsevi, sp. n.; 10-13) B. testacea; 6, 10) antenna, female; 7, 13) aedeagus, lateral aspect; 8, 11) theca of hypandrium, right lateral aspect; 9, 12) apical part of theca of hypandrium, left lateral aspect. Brachyopa cineria Wahlberg, 1844 Fig. 5
Zimina, 1981b: 31; Violovitsh, 1983: 100; Peck, 1988: 132.

MATERIAL. Khabarovskii krai: 2, 3, Komsomolsk-na-Amure, Silinskiy Park, 20,21.V 1986, 18.V 1993 (V. Mutin); Amurskaya oblast': 1, Zeya city, 24.VI 1982 (A. Ozerov); Primorskii krai: 1, 30 km N Ternei, 1.VI 1982 (V. 5


Mutin); 2, Vladivostok, Sedanka, 14.V 1983 (V. Mutin); 1, Luk'yanovka, 15.V 1981 (V. Mutin); 2, Anisimovka, 23.V 1986 (V. Makarkin). DISTRIBUTION. Russia: Amurskaya oblast', Khabarovskii krai, Primorskii krai; Siberia, European part. - Europe. Brachyopa dorsata Zetterstedt, 1837
Zimina, 1968: 67; 1981b: 31; Stackelberg, 1970: 39; Violovitsh, 1983: 99; Mutin, 1983a: 89; Peck, 1988: 132.

MATERIAL. Khabarovskii krai: 1, Gur River, larvae under bark of Populus tremula, 16.IX 1975 (Danilevskiy); 1, environs of Khabarovsk, larvae under bark of Populus tremula, 3.V 1975 (Zaitsev); 27, 14, Pivan village, 20.V 1979, 20.V 1988, 27.V 1990, 28-30.V 1991, 22.V, 7.VI 1992, 3.VI 1993, 25.VI 1994 (V. Mutin); 30, 5, Myaochan Range, Gornyi, 9.V 1983, 28.V, 8.VI 1988, 1.VI 1989, 31.V 1990, 6.VII 1991, 1.VII 1993 (V. Mutin); 13, 9, the mouth of Gorin River, 10,13.V 1984, 16.V 1985, 26,28.V 1986 (V. Mutin); 1, 45 km below Komsomolsk-na-Amure, 14.V 1986 (V. Mutin); 1, Komsomolsk-na-Amure, Silinskii Park, 14.V 1986 (V. Mutin); 2, environs of Komsomolsk-na-Amure, Snezhinka, 31.V 1989 (V. Mutin); 1; Amurskaya oblast': Zeya city, 20.VI. 1981 (A. Ozerov); Primorskii krai: 2, 30 km N Ternei, 31.V, 3.VI 1982 (V. Mutin); 4, Tigrovaya, 10-12.VI 1927 (Stackelberg). NOTES. The specimens (1 and 3) recorded as B. dorsata by L.V. Zimina (1972) belong to B. zhelochovtsevi sp. n. DISTRIBUTION. Russia: Primorskii krai, Khabarovskii krai, Amurskaya oblast', Siberia, European part. - Werstern Europe. Brachyopa maritima Violovitsh, 1980
Violovitsh, 1980: 125; 1983: 99; Mutin, 1984: 103; Peck, 1988: 132.

MATERIAL. Khabarovskii krai: 1, Gorin River, Zolotoy, 4.VI 1997 (V. Mutin); Primorskii krai: 4, Bol'shaya Ussurka River, Krutoi Yar, 17-20.VI 1995 (V. Mutin); 1, Kamenushka, 12,16.VI 1981 (V. Mutin); 5, 2, the same locality, 29.V - 3.VI 1989 (A. Shatalkin); 2, Vladivostok, Sedanka, 14.V. 1983 (V. Mutin); 1, Tigrovaya, 11.VI 1927 (Stackelberg); Sakhalinskaya oblast': 1 (holotype), Iturup I., Kurilsk, 27.VI 1957 (Violovitsh). DISTRIBUTION. Russia: Khabarovskii krai, Primorskii krai, Sakhalinskaya oblast'(Kuril Islands: Iturup, Kunashir). Brachyopa ornamentosa Violovitsh, 1977
Violovitsh, 1977: 81, 1983: 100; Thompson, 1980: 211; Peck, 1988: 132.

MATERIAL. Primorskii krai: 1, Pos'et, 1.V 1978 (Barkalov); Khabarovskii krai: 15, 14, Komsomolsk-na-Amure, Silinskii Park, 8.V 1985, 20.V 1986, 6


17,23.V 1987, 17.V 1988, 5, 9.V 1990 (V. Mutin); 1, Myaochan Range, Gornyi, 28.V 1988 (V. Mutin). DISTRIBUTION. Russia: Khabarovskii krai, Primorskii krai. - East China. Brachyopa panzeri Goffe, 1945
Mutin, 1983b: 101.

MATERIAL. Amurskaya oblast': 15, , Klimoutsy, 30.V- 4.VI 1958 (Borisova, Zinov'ev); Primorskii krai: 3, 4, 30 km N Ternei, 30.V, 11,13.VI, 3,5.VII 1982 (V. Mutin); 1, Luk'yanovka, 15.V 1981 (V. Mutin); 1, Kamenushka, 13.VI 1981 (V. Mutin); 1, Anisimovka, 17.VII 1974 (Yu. Berezantsev); 1, Vinogradovka, 24.VI 1927 (Stackelberg); 1, the same locality, 10.VI 1929 (Filip'ev, D'yakonov). DISTRIBUTION. Russia: Amurskaya oblast' (new record), Primorskii krai; European part. - Europe. Brachyopa pivanica Mutin, 1984
Mutin, 1984: 102.

MATERIAL. Khabarovskii krai: 4 (including the holotype and paratype), Pivan', 6.VI 1979, 27.V 1990, 7.VI 1995 (V. Mutin); 2, 1, Myaochan Range, Gornyi, 8.VI 1988, 1.VII 1993, 19.VI 1997 (V. Mutin); Amurskaya oblast': 3, 2, Zeya city, 9-30.VI. 1978; 11.VII 1981 (A. Ozerov, A. Shatalkin); Primorskii krai: 1, 25 km N Ternei, 10.VII 1982 (V. Mutin); DISTRIBUTION. Russia: Primorskii krai (new record), Khabarovskii krai, Amurskaya oblast', Transbaicalia, Jakutia. Brachyopa testacea (FallИn, 1817) Figs 10-13
Zimina, 1981a: 157; Violovitsh, 1983: 99; Peck, 1988: 113.

MATERIAL. Khabarovskii krai: 10, 2, Myaochan Range, Gornyi, 28.V 1988, 30.VI 1987, 6.VII 1991, 14-22.VII 1994 (V. Mutin); 1, the same locality, 13.VII 1996 (D. Gritskevich); 3, the mouth of Gorin River, Kholodnyi Stream, 30.V 1984, 10.VI 1985 (V. Mutin); Amurskaya oblast': 1, Zeya town, 4.VII 1981 (A. Shatalkin); Primorskii krai: 1, 30 km N Ternei, 3.V 1982 (V. Mutin); DISTRIBUTION. Russia: ?Chukotka, Khabarovskii krai, Amurskaya oblast', Primorskii krai, Siberia, European part. - Mongolia, Europe. 7


Brachyopa violovitshi Mutin, 1985
Mutin, 1985: 85.

MATERIAL. Khabarovskii krai: 84, Komsomolsk-na-Amure, Silinskii Park, 8,11.V 1985, 20,22.V 1986, 17.V 1988, 9.V 1990, 23.V 1992, 18.V 1993, 25.V1994 (V. Mutin); 2, Pivan', 27.V 1990; 3, Myaochan Range, Gornyi, 28.V, 1.VI 1988 (V. Mutin); Primorskii krai: 1 (holotype), 30 km N Ternei, 27.V 1982 (V. Mutin); 1 (paratype), Kavalerovo, 22.V 1982 (V. Mutin); 1 (paratype), Vladivostok, Sedanka, 5.V 1982 (V. Mutin); 1 (paratype), Kamenushka, 14.VI 1981 (V. Mutin); 1, Vinogradovka, 18.V 1929 (Filip'ev, D'yakonov). DISTRIBUTION. Russia: Khabarovskii krai, Primorskii krai. Brachyopa vittata Zetterstedt, 1843
Violovitsh, 1983: 99; Peck, 1988: 133.

MATERIAL. Khabarovskii krai: 16, 11, Myaochan Range, Gornyi, 25.VI 1982, 25.VI, 7.VII 1986, 30.VI 1987; 8.VI 1988; 1.VII 1993; 14-23.VII 1994 (V. Mutin); 1, the mouth of Gorin River, 16.VI 1984 (V. Mutin); Primorskii krai: 1, Livadiyskii Range, Mt. Krinichnaya, 1200 m, 19.VI 1982 (V. Mutin); 1, Mt. Ol'khovaya, 1600 m, 8.VIII 1986 (V. Makarkin). DISTRIBUTION. Russia: Khabarovskii krai, Primorskii krai, ?Sakhalinskaya oblast' (Kuril Islands: Kunashir), South Siberia, European part. - Europe. REMARKS. I did not find any specimens of B. vittata from Kuril Islands in the Russian collections. Brachyopa bicolor (FallИn, 1817)
Zimina, 1968: 66; Violovitsh, 1983: 99; Peck, 1988: 132.

DISRTIBUTION. Russia: ? Primorskii krai, south of European part. Caucasus. REMARKS. All above-mentioned authors don't listed specimens of B. from Primorskii krai or other parts of Russian Far East. In collections I did any specimens from this region. Probably B. bicolor not distributed in the Far East. ACKNOWLEDGEMENTS

Europe, bicolor not find Russian

I would like to express my gratitude to Dr A.L. Ozerov and Dr. A.I. Shatalkin for opportunity to study collections of the Zoological Museum of Moscow State University. I thank Dr. A.V. Barkalov (Institute of Systematics and Animal Ecology, Novosibirsk) and Dr. S.Yu. Kuznetsov (Zoological Institute, St.Petersburg) for their help as curators of syrphid collections in those institutions. 8


REFERENCES
Mutin, V.A. 1983a. [Fauna and ecology of hover flies (Diptera, Syrphidae) as pollinators of some Angiosperms of the Lower Amurland]. - In: Soboleva, R.G. (Ed.). Obzor otdel'nykh otryadov nasekomykh Dal'nego Vostoka. Vladivostok: 86-99. (In Russian). Mutin, V.A. 1983b. [Hover-flies (Diptera, Syrphidae) of anthophilous complexs of the Southern Primorskii krai]. - In: Soboleva, R.G. (Ed.). Obzor otdel'nykh otryadov nasekomykh Dal'nego Vostoka. Vladivostok: 100-109. (In Russian). Mutin , V.A. 1984. [New and little-known species of hover-flies (Diptera, Syrphidae) from the South of the Far-East]. - In: Tsherepanov, A.I. (Ed.) Novye i maloizvestnye vidy fauny Sibiri. Vyp. 17. Chlenistonogie i gel'minty. Novosibirsk: 100-106. (In Russian, with English summary). Mutin, V.A. 1985. [New data on hover-flies (Diptera, Syrphidae) of Far East]. - In: Taxonomiya i ekologiya chlenistonogikh Dal'nego Vostoka. Vladivostok: 85-99.(In Russian). Peck, L.V. 1988. Family Syrphidae. In: Soos, A. (Ed.). Catalogue of Palaearctic Diptera. Budapest. Pp. 11-230. Stackelberg, A.A. 1970. [Fam. Syrphidae]. - In: Opredelitel' nasekomykh evropeiskoi chasti SSSR. Tom. V. Vyp. 2. Moscow-Leningrad: 11-96. (In Russian). Thompson, Ch. 1980. The problem of old names as illustrated by Brachyopa "conica Panzer" with a synopsis of Palaearctic Brachyopa Meigen (Diptera: Syrphidae). - Ent. scand., 11: 209-216. Violovitsh, N.A. 1977. [Some new Palaearctic species of hover flies (Diptera, Syrphidae)]. In: Cherepanov, A.I. (Ed.). Novye i maloizvestnye vidy fauny Sibiri. Vyp. 11. Taksony fauny Sibiri. Novosibirsk: 68-84.(In Russian, with English summary). Violovitsh, N.A. 1980. [New species of flower flies (Diptera, Syrphidae) of the Palaearctic fauna]. - In: Cherepanov, A.I. (Ed.). Novye i maloizvestnye vidy fauny Sibiri. Vyp. 14. Sistematika i ekologiya zhivotnykh. Novosibirsk: 124-131.(In Russian, with English summary). Violovitsh, N.A. 1983. Sirphidy Sibiri (Diptera, Syrphidae). Opredelitel. Novosibirsk: 241 pp. (In Russian). Zimina, L.V. 1968. [To dipterofauna of Estern Siberia, Syrphidae and Conopidae]. - In: Zhelokhovtsev, A.N. (Ed.). - In: Trudy Zoologicheskogo muzeya Moskovskogo universiteta. Tom 11. Issledovaniya po faune Sovetskogo Soyuza (nasekomye). Moscow: 57-78. (In Russian). Zimina, L.V. 1972. [Syrphids (Diptera, Syrphidae) of Magadanskaya oblast']. - Bulleten' Moskovskogo obstshestva ispytatelei prirody 77 (1): 37-45.(In Russian, with English summary). Zimina, L.V. 1981a. [Rare and interesting Syrphidae (Diptera) in the collection of the Zoological Museum of MSU]. - In: Trudy Zoologicheskogo muzeya Moskovskogo universiteta. Tom 19. Nasekomye (Issledovaniya po faune Sovetskogo Soyuza). Moscow: 150-170. (In Russian). Zimina, L.V. 1981b. [Syrphids (Diptera, Syrphidae) of north Amur district]. - In: Ekologofaunisticheskie issledovaniya. Biologicheskie resursy territorii stroitel'stva zony BAM. Moscow: 27-38. (In Russian).

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SHORT COMMUNICATION
D. I. Grit sk ev ich. HOVER-F LI ES (D IPTERA: SYR P HIDAE) IN ANTHOPHILOUS COMPLEXES OF PLANTS OF MYAOCHAN RANGE, KHABAROVSKII KRAI. - Far Eastern Entomologist. 1998. N 65: 10-14. .. . - (Diptera: Syrphidae) , . // . 1998. N 65. C. 10-14.
Two hundred four species in 52 genera of hover-flies are known from Myaochan Range (Khabarovskii krai) [5]. There are four phenological groups of syrphids in Lower Amur region [4]. There is a strict dependence of hover-flies species composition belonging to anthophilous complex of plants from their blossoming. The material was collected during 1994-1996 using a standard entomological net. The degree of difference of anthophilous complexes was studied by means of the method of multidimensional scaling (degree of difference - euclidean distances) [6]. Multidimensional scaling allows to present a sharing of the objects in the coordinate space with the small quantity of dimensions graphically. One hundred five species of hover-flies visit seven most abundant plants (Table 1). Anthophilous complexes of plants of Myaochan Range forms three well isolated groups (Fig. 1). The first one includes plants with peak of blossoming in the middle of June (Rhododendron

Fig.1. Position of the anthophilous complexes of plants in the reduced space of the three coordinates, based on the scaling of matrix euclidean distances.

10


Table 1. Syrphids in anthophilous complexes of plants (I - Weigela middendorffiana; II - Rhododendron aureum; III - Trollius riederanus; IV - Aruncus asiaticus; V - Heracleum moellendorffii; VI - Chamerion angustifolium; VII - Saussurea dubia) in Myaochan Range Species I Blera eoa (Stack.) B. judini Bark. Brachyopa dorsata Ztt. B. vittata Ztt. Callicera aenea (F.) Chalcosyrphus nitidus (Portsch.) Ch. rufipes (Lw.) Ch. tuberculifemur (Stack.) Cheilosia impressa Lw. Ch. longula (Ztt.) Ch. motodomariensis Mats. Ch. pagana (Mg.) Ch. pallipes Lw. Ch. primoriensis Bark. Ch. reniformis (Hellen) Ch. scutellata (Fallen) Ch. sichotana (Stack.) Ch. velutina Lw. Chrysotoxum fasciatum (Mull.) Ch. fasciolatum (DeGeeer) Ch. sapporense Mats. Criorhina aino (Stack.) C. brevipila Lw. C. ussuriana (Stack.) Dasysyrphus bilineatus (Mats.) D. lunulatus (Mg.) D. nigricornis (Verr.) D. tricinctus (Fallen) D. venustus Mg. D. zinchencoi Mut. et Bark. Didea alneti (Fallen) D. fasciata (Mcq.) Epistrophe grossulariae (Mg.) Episyrphus balteatus (DeGeer) Eriozona (Megasyrphus) erratica (L.) E. syrphoides (Fallen) Eristalis alpina (Pz.) E. cerialis F.
3.82 1.53 0.76 3.82 0.76 0.76 0.76 1.53 1.53 1.53 3.05 -

II

Plants (share of syrphids, %) III IV V VI
0.60 0.20 0.20 0.99 1.79 10.12 0.79 0.60 0.20 11.71 0.20 0.20 0.20 0.79 1.59 0.20 0.40 0.64 0.64 2.55 1.91 0.64 0.64 0.64 5.45 0.71 8.53 0.95 0.47 0.24 0.24 0.24 0.47 0.24 0.24 0.24 0.95 8.33 2.78 5.56 2.78 -

VII
2.56 2.56 2.56 5.13 -

0.26 0.26 0.26 0.26 1.32 11.61 0.26 0.26 1.06 0.26 0.53 0.26 0.53 0.79 1.06 0.53 0.26 1.85 1.06 0.26 0.26

11


Table 1 (continued)
Species I E. interrupta (Poda) E. rabida Viol. E. rossica Stack. E. rupium F. E. vitripennis Strobl Eupeodes(Lapposyrphus) lapponicus (Ztt.) E. latilunulatus (Collin) E. lundbecki (Soot-Ryen) E. luniger (Mg.) E. nitens (Ztt.) Helophilus affinis Wahlb. Lejota korsakovi (Stack.) Leucozona (Ischyrosyrphus) glaucius (L.) L. (I.) laternarius (Mull.) L. lucorum (L.) Melangyna barbifrons (Fallen) M. coei Niel. M. compositarum (Verr.) M. lasiophthalma (Ztt.) M. motodomariensis (Mats.) M. quadrimaculata (Verr.) Melanostoma boreomontanum Mut. M. mellinum (L.) M. scalare (F.) Meliscaeva cinctella (Ztt.) Neocnemodon vitripennis (Mg.) Orthonevra stackelbergi Thomp.et T.-Ped. Parasyrphus annulatus (Ztt.) P. lineolus (Ztt.) P. macularis (Ztt.) P. malinellus (Collin) P. nigritarsis (Ztt.) P. punctulatus (Verr.) P. tarsatus (Ztt.) Pipiza bimaculata Mg. P. quadrimaculata (Pz.) P. signata Mg. Platycheirus albimanus (F.) P. angustatus (Ztt.) P. clypeatus (Mg.) P. coerulescens Will. P. complicatus (Beck.) P. europaeus Goeld.Maib.et Speig
1.53 1.53 0.76 2.29 1.53 4.58 0.76 0.76 1.53 12.21 6.11 3.82 0.76 3.05 0.76 0.76 2.29 3.82 -

Plants (share of syrphids, %) II III IV V VI
0.53 0.53 0.26 5.54 0.26 0.53 0.26 0.79 1.06 0.26 0.79 2.11 0.79 4.22 0.53 7.12 3.69 2.64 0.26 0.53 0.79 2.37 0.53 1.32 0.26 1.06 0.53 0.79 0.40 0.40 6.15 0.20 0.20 0.20 0.20 0.60 1.19 0.60 0.40 3.57 1.19 2.78 1.19 1.79 1.19 12.50 7.14 0.60 0.60 0.60 0.99 0.20 1.39 0.60 0.20 0.20 0.64 1.27 2.55 1.27 0.64 0.64 0.64 5.10 2.55 20.38 1.27 2.55 0.64 3.82 1.27 1.27 0.47 1.66 1.66 0.71 0.71 15.88 1.42 0.95 40.05 0.24 0.47 0.24 0.24 1.18 0.24 8.33 5.56 2.78 2.78 2.78 -

VII
5.13 1.28 8.97 10.26 -

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Table 1 (continued)
Species I P. peltatus (Mg.) P. scutatus (Mg.) P. urakawensis (Mats.) Pseudopocota stackelbergi (Viol.) Scaeva komabensis (Mats.) Sericomyia lappona (L.) Sphaerophoria chongjini Bank. Sphegina (Asiosphegina) sibirica Stack. Spheginoides obscurus Szil. Syrphus ribesii (L.) Syrphus sp. S. torvus O.-S. S. vitripennis Mg. Temnostoma apiforme (F.) T. vespiforme (L.) Volucella bombylans (L.) Xanthandrus comtus (Harr.) Xylota atricoloris Mut. X. coeruleiventris (Ztt.) X. coquilletti H.-B. X. ignava (Pz.) X. pseudoignava Mut. X. sibirica Lw. X. silvicola Mut. Nubmer of species
0.76 2.29 0.76 10.69 4.58 6.87 0.76 0.76 3.05 0.76 -

Plants (share of syrphids, %) II III IV V VI
0.53 0.26 0.26 5.28 15.04 8.18 0.26 0.53 0.26 1.58 0.53 4.49 1.98 0.20 0.20 0.20 5.16 0.20 7.94 1.79 0.20 0.20 0.79 0.20 1.98 0.20 0.64 0.64 0.64 1.91 31.85 3.18 0.64 1.27 1.27 1.27 1.91 0.64 0.95 0.24 3.79 1.42 1.90 1.66 2.37 0.95 0.71 0.24 0.47 0.24 2.78 5.56 2.78 2.78 5.56 27.78 11.11 -

VII
1.28 2.56 53.85 3.85 -

39

60

60

35

40

16

12

aureum, Trollius riederanus, Weigela middendorffiana), the second group - in July (Heracleum moellendorffii, Aruncus asiaticus), in the third one - in the middle and end of August (Chamerion angustifolium, Saussurea dubia). Early-summer phenological group of hover-flies mainly visit the plants which are blossoming in June [4]. Anthophilous complexes of Rhododendron aureum and Trollius riederanus are most similar because of abundance of pollinophagous hover-flies from genera Syrphus, Parasyrphus, Platycheirus, Cheilosia [3]. Syrphids of these genera visit an opened actinomorphous, rich of pollen flowers. A complex of Weigela middendorffiana has more differences from previous ones. There are many large nectarophagous syrphids (Criorhina, Pseudopocota, Eriozona, Didea) among visitors of Weigela flowers with tubular corolla. The larvae of some nectarophagous dendrobionts syrphids are saproxylophagous [2], for example Criorhina, Mallota, Temnostoma, Chalcosyrphus. There are a lot of species of hover-flies from the late-summer phenological group (Cheilosia motodomariensis, Ch. impressa, Ch. velutina, Melangyna compositarum, M. motodomariensis) in anthophilous complexes of flowering in July plants (Heracleum moellendorffii, Aruncus asiaticus). Cheilosia motodomariensis, Ch. impressa and Leucozona glaucius are feeding mainly on H. moellendorffii [1]. These syrphids are rare on A.

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asiaticus. The dominant species on A. asiaticus is Sphegina sibirica (31,8%). Therefore distinct differences in anthophilous complexes of H. moellendorffii and A. asiaticus are observed (Fig. 1). In anthophilous complexes of flowering in August plants the hover-flies are not numerous. Almost all these syrphids are dominants on the Myaochan Range during the summer season (Syrphus torvus, Eupeodes lapponicus, Parasyrphus lineolus, P. annulatus, Platycheirus albimanus, Cheilosia longula). Therefore the similarity of anthophilous complexes of Chamerion and Saussurea is the highest (Fig. 1). 1. Bagathsanova, A.K. 1990. [Fauna and ecology of hover-flies (Diptera; Syrphidae) of Yacutia]. Yakutsk: 162 pp. (In Russian). 2. Krivosheina, N.P. & Zaitsev, A.I. 1989. [Phylogenesis and evolutionary ecology of dipterous insects]. - Itogi nauki i tekhniki. Ser. Entomologiya. Moscow. 9: 92-95. (In Russian). 3. Mutin, V.A. 1987. [rophic relationship of imago of syrphids (Diptera; Syrphidae) with floral plants]. - In: Dvukrylye nasekomye: sistematika, morphologiya, ekologiya. Leningrad: 77-79. (In Russian). 4. Mutin, V.A. 1992. [Phenological aspects of fauna of hover-flies (Diptera; Syrphidae) of the south of Far East]. - In: Systematika, zoogeografiya i kariologiya dvukrylykh nasekomykh (Insecta; Diptera). St. Petersburg: 119-121. (In Russian). 5. Mutin, V.A. & Gritskevich, D.I. 1998. [Ecology-faunistic review of hover-flies (Diptera; Syrphidae) of Lower Amur region.] - In: A.I. Kurentzov's Annual Memorial Meetings. Issue VIII. Vladivostok: 71-86. (In Russian). 6. Terjokhina, A.Yu. 1986. [Data analyses by multidimensional scaling methods]. Moscow: 168 pp. (In Russian). Author's address: Department of Zoology, Komsomolsk-na-Amure State Pedagogical Institute, 681000, Russia

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SHORT COMMUNICATION
D.S. Aristov. A NEW SPECIES OF THE PERMIAN GENUS KUNGURMICA (GRYLLOBLATTIDA: PERMEMBIIDAE). - Far Eastern Entomologist. 1998. N 65 : 15-16. .. . Kungurmica (Grylloblattida: Permembiidae). // . 1998. N 65. C. 15-16.
In 1928 R.J. Tillyard described a very small insect Permembia delicatula from Lower Permian of North America in the family Delopteridae [3]. Later a new family Permembiidae was erected in order Psocoptera [4]. Now Permembiidae is included in the order Grylloblattida [2]. Kungurmica tchekardensis was described recently from Lower Permian of Ural [1]. New species of Kungurmica is found in the Upper Permian of North Russia. I am grateful to Dr. V.G. Novokshonov for support in preparation of manuscript. Family Permembiidae Tillyard, 1937 Genus Kungurmica Novokshonov, 1998 Kungurmica sojanensis Aristov, sp. n. Fig. 1 MATERIAL. Holotype: imprint and counter-imprint of body, specimen N 3353/233; Russia, Arkhangelsk region, right bank of Sojana River about 57 km from its mouth, IvaGora; Upper Permian, Kazanian Stage; in collection of Paleontological Institute, Moscow.

Fig. 1. Kungurmica sojanensis sp. n., holotype, spec. N 3353/233. Scale bar ­ 1 mm.

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DIAGNOSIS. New species due to bed preservation of insect is formaly included to the genus Kungurmica based on the small size, the shape of head and antennae. However K. sojanensis is distinguished from single speciment of K. tchekardensis by short pronotum and more longer legs. But the main reason for erecting of new species is the difference in age of both species. DESCRIPTION. Small insect, lengh of body 3.5 mm. Body elongate. Head prognathous, robust, with acute mandibles projecting far forwards. Antennae moniliform, with 10 pear-like segments. Eyes large. Pronotum very short, mesonotum trapezoidal, larger than metanotum. Legs probably long, hind legs reaching to wing apex. Wings are quitely not keeping, lengh of forewing about 2.6 mm. Abdomen with 7 visible segments, reaching to wing apex or slightly shorter. 1. Novokshonov, V.G. 1998. New fossil insects (Insecta; Grylloblattida, Caloneurida, Hypoperlida, Ordinis incertis ) from the Kungurian Beds of the Middle Urals. - Paleontological Journal 4: 41-46. (In Russian). 2. Storozhenko, S.Yu. 1997. Classification of order Grylloblattida (Insecta), with description of new faxa. - Far Eastern Entomologist 42: 1-20. 3.Tillyard, R.J. 1928. Kansas Permian insects. Part 12. The family Delopteridae, with a discussion of its ordinal position. - Amer. J. Sci., 16: 469-484. 4. Tillyadr, R.J. 1937. Kansas Permian insects. Part 17. The order Megasecoptera and additions to the Paleodictyoptera, Odonata, Protoperlaria, Copeognata and Neuroptera. Amer. J. Sci., 33: 81-100. Author's address: Department of Biology, Perm State University, Perm, 614600, Russia

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