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Studia dipterologica 3 (1 996) Heft 1

KRIVOSHEINAKRIVOSHEINA:Raineria... Russia... (97-100) & ...

The species from the genus Rainieria RONDANI (Diptera, Micropezidae) of Russia including the description of a new species
[Die Arten der Gattung Rainieria RONDANI (Diptera, Micropezidae) aus RuDland nebst der Beschreibung einer neuen Art] by Marina G. KRIVOSHEINA and Nina P. KRIVOSHEINA
Moscow (Russia) Moscow (Russia)

Abstract

The wide distribution ofR. latifrons (LOEW) Europe to the Kuril islands (Kunashir) from is proved. A new species, Rainieria hennigi spec. nov. from Kunashir, is described. It was apparently mentioned before by HENNIG (1938) under the name of R. latifrons. The data on the distribution of R. latifrons in Japan need confirmation. A key to the is species of Rainieria RONDANIgiven. Diptera, Micropezidae, Rainieria RONDANI, species, Russia, distribution, key to new the species. Die weite Verbreitung von R. latifrons (LOEW) zu den Kurilen (Insel Kunaschir) bis wird bestatigt. Es wird eine neue Art Raineria hennigi spec. nov. von der Insel Kunaschir beschrieben. HENNIG (1938) erwahnte diese Art unter der Bezeichnung R. latifrons. Die Angaben uber die Verbreitung von R. latifrons in Japan miissen iiberpruft werden. Es wird eine Bestimmungstabelle Wr die Arten der Gattung Rainieria RONDANI gegeben. Diptera, Micropezidae, Rainieria RONDANI, neue Art, RuRland, Verbreitung, Bestimmungstabelle der Arten.

Key words

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Zusammenfassung

Stichworter

Introduction
Three species from the genus Rainieria RONDANI distributed in Palaearctic Region: R. are 1935), R. calceata (FALLEN, 1820) and R. latijirons (LOEW, 1870). The boninensis (HENNIG, representatives of the genus are characterized by the presence of well developed postocellar setae and massive shining clypeus projecting beyond the mouth cavity. Raineria latifrons is the most common and widely distributed species. It was registered from Hungary and Romania to Japan (STACKELBERG Sobs 1984). Raineria calceata is known from vari1970; ous areas of Europe, as well as from southern Russia, and R. boninensis from Japan. HENNIG (1938) gave short diagnostical descriptions of the 3 palaearctic species. The data on R. latifrons were based on the material from Hokkaido and the northern part of Honshu. The author might have adopted CZERNY'S opinion (1930) that it was the only species distributed in Japan, and determined it as R. latifrons only because of the place of collecting. We examined specimens of R. calceata and R. latifrons, kept in the collections of the Zoological Museum of the Moscow Lomonosov State University [ZMUM] and A.N. SEVERTZOV Institute of Ecology and Evolution (IEE, Moscow). The material included large series of imagoes ofR. latifrons from the territory of the former USSR from the Karpathians to the Far East. As a result of this it was shown that the part of the material of R. latifrons


KRIVOSHEINAKRIVOSHEINA:Raineria... Russia... (97-100) & ...

Studia dipterologica 3 (1 996) Heft 1

from Kunashir Island proved to belong to a new species, which is very close to the previousely known one. Jurging by the figures, the specimens of R. latifrons mentioned by HENNIG (1938) from Hokkaido and Honshu belong to the same new species. Description of species

Rainieria hennigi spec. nov. (Figs. 1-3)
Diagnosis Median frontal stripe formed by velvety-grey pollen relatively narrow, 2 times as wide as the orbital plate. The 3rd antennal segment and arista completely dark. Preapical light spot of the middle femora 2 times as long as the dark apical part (dorsal view). Median band of wing indistinct, the apex of R, light. Epandrium (Fig. 1) relatively short, narrowing apically. Sickle-like appendages of the copulative fork with black thick setae of various length, contacting at median line. Description Male: body length 7,5 mm, length of wing - 6 mm. Body brown. Head round, brown with strongly projecting shining black clypeus. Eyes oval, bare. Frons wider than the eye and slightly exceeding its length from anterior ocellus to the 1unula.Velvety-grey pollen forms a spindle-like median frontal stripe. In front of the ocellus it is twice as wide as the orbital plate. 1 orb s, 2 fr s. Orbital setae are closer to frontal setae, than to vertical setae. The lower half of postcranium with silvery V-shaped spot, formed by dense short pubescence, and several long setae, the 4-5 of which forming a row. Narrow parafacial and genae with yellowish-silvery pubescence. Antennae and arista dark-brown, almost black, the 3rd antennal segment in dense short greyish pubescence. Palpae black. Thorax brown with grey pollen. Mesonotum with 2 npl s and 2 spa1 s. 2 dc in front of scutellum are long. Anepimeron with 2 rows of long setae on posterior margin. Scutellum broadly rounded with 3 pairs of short setae. Postnotum with bluish pollen without pubescence. Legs: coxa elongated, brown, fore coxa lighter, yellowish at apex but darkened at exterior surface. All femora light in base. Middle and hind femora with yellow spot near apex, the latter of the hind femora (dorsal view) being elongated, 2 times as long as the dark apex. Tibia dark. Fore tarsi snow-white, middle and hind tarsi brown but hind metatarsus light yellowish with dense light short pubescence. Halteres dark. Coloration pattern of wing indistinct. Median dark band is situated basally to R,, slightly projecting beyond rm. Abdomen lighter in base and darker, almost black at apex, with bluish pollen on apical half. First abdominal segment with long erected light setae, the following segments with short brown lying setae. Genitalia: epandrium relatively short, slightly narrowing apically (lateral view). Copulative fork of the 5th segment with long sickle-like apical appendages which are about as long as or longer than the stem of the fork. Sickle-like appendages with long dense black thick setae (Fig. 2), situated on their inner margin and brought together near the median line. Exterior margin with long black erected setae. Female Resembling male but the next characters. Light spot in front of the apex of middle femora is as long as the dark apical part. Light spot in front of the apex of the hind femora is no more than 1,5 times as long as the dark apical part. Biology: imagoes of the species were reared from the larvae taken under the bark of birch (Betula) and fir (Abies). Distribution: Far East: Russia, Kurile, Kunashir; Japan: Hokkaido, Honshu (Hondo).


Studia dipterologica 3 (1 996) Heft 1

KRIVOSHEINAKRIVOSHEINA:Raineria... & ...

Russia... (97-100)

Material: Holotype 6: Russia, Kurile, Kunashir, volkano Mendeleeva, 28 June 1977 (leg. A.I. ZAITZEV); Paratypes: Russia, Kurile, Kunashir, volkano Mendeleeva, 18 April 1977: 16, I?; 28 June 1977: 16, 1 0; Russia, Kurile, Kunashir, Tretjakovo, 17 July 1977: 26 6 (leg. A.I. ZAITZEV) [ZMUM].

Rainieria latifrons (LOEW, 1870), (Figs. 4-7)

Diagnosis Ma1e:median frontal stripe wide, 2,5-3,O times as wide as the orbital plate in front of the ocellar triangle. The 3rd antenna1 segment and arista red-yellow in base. Face with black band below antennae, this band absent in new species. Preapical light spot of the middle and hind femora is about as long as the dark apical part. Median band of wing wide, apex of R, darkened, the band reaching posterior margin of wing. Dark apical spot large. Epandrium relatively long (Fig. 5), not narrowed apically. Sickle-like appendages of the copulative fork with short spines of equal size on the inner margin. Biology: larvae breed in friable light wood of decidious: beech (Fagus), hornbeam (Rosa), birch Betula), oak (Quercus), asp (Populus), lime-tree (Tilia), and Chosenia. Distribution: wide Palaearctic, Far East, including Kunashir (Kurile).

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Figs. 1-7: Genital structures of Raineria hennigi spec. nov. (1-3) and Raineria latifrons (Lo~w,1870)(4-7); - 1: female, tip of abdomen; 2: male copulative fork, exterior view and - 3: lateral view; - 4: male abdomen, lateral view (x32); - 5: female, tip of abdomen; - 6: male copulative fork, exterior view and - 7: lateral view. Figs. 1-3; 5-7 ~ 6 4 .

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Material: Russia, Kaluzhskaya Area, Kirejkovo, 2 June 1992: 16, 29 ? (leg. A.I. ZAITZEV); Russia, Krasnodarsky Area, Ubinskaya , 9 July 1970: 26 6 , 29 ? , 20 August 1970: 16, 1 ? (leg. N.P. KRIVOSHEINA); Russia, Amurskaya Area, Kundur, 21 May 1975: 16; 24 May 1975: 16 ; 2 June 1975: 366, 10 (leg. A.I. ZAITZEV); Russia, Amurskaya Area, Zeja, 12 June 1981: 26 6 (leg. A.L. OZEROV); Russia, Khabarovsky Kraj, Khabarovsk, 3 July 1975: 16 (leg. B.M. MAMAEV); Russia, Khabarovsky Kraj, Khabarovsk, 18 April 1976: 23 Russia, 66 8, May 1976: 16 (leg. N.P. KRIVOSHEINA). Primorskij Kraj, 32 km SE Ussurijsk, 15 September


KRIVOSHEINA & KRIVOSHEINA: ... Raineria... Russia... (97-100)

Studia dipterologica 3 (1 996) Heft 1

1964: 1 0, 7 May 1967: 1 0, 7 June 1968: 1 0, 17 May 1969: 1 0 (leg. N.P. KRIVOSHEINA); Russia, Primorskij Kraj, 32 km SE Ussurijsk, 26 August 1987: 18 (leg. A.V. ANTROPOV); Russia, Primorskij Kraj, 32 km SE Ussurijsk, 6 August 1948: 18 (leg. V.V. GUSSAKOVSKY); Russia, Primorskij Kraj, Preserve ,,Kedrovaya Pad", 15-22 August 1985: 30 0 (leg. N.P. KRIVOSHEINA); Russia, Primorskij Kraj, Lazo, 14 July 1979: 10, 26 July 1979: 18 (leg. N.P. KRIVOSHEINA); Russia, Kurile, Kunashir, Tretjakovo, 21 July 1985: 16 (leg. S. CHURKIN).

Key to the species of Rainieria RONDANI 1

2

All femora black at base. Lunule with long light hairs. Median frontal stripe no 1820) more than 2 times as wide as the orbital plate .......... R. calceata (FALLEN, All femora or at least fore femora light at base ..................................................... 2 Epandrium elongated, 2,s-3 times as long as high (lateral view). Sickle-like appendages of the copulative fork more than 2 times as short as the stem of the fork ........................................................................... R. boninensis (HENNIG, 1935) Epandrium short, no more than 1,s times as long as high. Sickle-like appendages of the copulative fork as long as or slightly shorter than the stem of the fork ... 3 Median frontal stripe at least 2,s times as wide as orbital plate. Sickle-like appendages of the copulative fork with short black spines of equal size on the inner margin ....................................................................R. Iatifrons (LOEW, 1870) Median frontal stripe no more then 2 times as wide as orbital plate. Sickle-like appendages of the copulative fork with long thick black setae of various size on the inner surface .................................................................... R. hennigi spec. nov.

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Acknowledgements
The research described in this publication was made possible in part by the joint Grant N MDA 300 from the International Science Foundation (J. So~os) Russian Government, as well as the Russian Foundation for and Fundamental Researches N 94- 1127 la.

Literature
STACKELBERG, (1970): Fam. Calobatidae (Trepidariidae). - In: STACKELBERG, &NARTSCHUK, (eds.): A.A. A.A. E.P. The key to the insects of the European part of the USSR 5(2): 112-114; Leningrad. CZERNY, (1930): 42 a. Tylidae. - In: LINDNER, (ed.): Die Fliegen der palaearktischen Region 5(1): 1-17; L. E. Stuttgart. W. HENNIG, (1938): Tyliden aus Japan. - Insecta matsumara 13: 1-14; Sapporo. Sob, A. (1984): Family Micropezidae. In: So6s, A. & PAPP, (eds.): Catalogue of Palaearctic Diptera 9: 19L. 24; Budapest.

Authors' address Dr. Marina.G. KRIVOSHEINA Prof. Dr. Nina P. KRIVOSHEINA Institute of Ecology and Evolution Russian Academy of Sciences 33 Leninsky Prospect 117071 Moscow Russia

The paper was received on 20 January 1996.