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SOROKINA: 29­39

Studia dipterologica 16 (2009) Heft 1/2 · ISSN 0945-3954

The genus Spilogona SCHNABL (Diptera: Muscidae) in north European Russia: new records and new species
[Die Gattung Spilogona SCHNABL (Diptera: Muscidae) im nordeuropÄischen Russland: neue Nachweise und neue Arten] by Vera S. SOROKINA
Novosibirsk (Russia)

Abstract

A list is given of the species of the genus Spilogona SCHNABL, 1911 (Muscidae) of the North European Territory of Russia. 39 species of Spilogona are listed, including two new species from the Archangel'skaya oblast': Spilogona vikhrevi spec. nov. and Spilogona archangelica spec. nov. Spilogona genualis HUCKETT, 1965 is newly recorded from Russia and the Palaearctic region. Spilogona norvegica (RINGDAHL, 1932) and Spilogona nitidicauda (SCHNABL, 1911) are newly recorded from the North European Territory of Russia. The male terminalia of the new species are figured. Muscidae, Spilogona, Palaearctic, European Russia, Archangel'sk region, Naryan-Mar, taxonomy, faunistics, new species, new records, keys

Key words

Zusammenfassung Die Liste der Arten aus der Gattung Spilogona SCHNABL, 1911 (Muscidae) im nordeuropÄischen Russland umfasst 39 Arten, einschlieúlich zweier neuer Arten aus dem Archangelsker Gebiet: Spilogona vikhrevi spec. nov. und Spilogona archangelica spec. nov. Spilogona genualis HUCKETT, 1965 ist neu fÝr Russland und die PalÄarktische Region. Spilogona norvegica (RINGDAHL, 1932) und Spilogona nitidicauda (SCHNABL, 1911) sind neu fÝr das nordeuropÄische Russland. Der mÄnnliche Genitalapparat der neuen Arten wird abgebildet. StichwÆrter Muscidae, Spilogona, PalÄarktis, europÄisches Russland, Archangelsker Gebiet, Naryan-Mar, Taxonomie, Faunistik, neue Arten, neue Nachweise, BestimmungsschlÝssel

Introduction Flies of the genus Spilogona SCHNABL, 1911 are widespread in the Holarctic region, but the distribution of the genus is uneven because most species prefer cold habitats. The genus is extremely speciose at high altitudes and arctic latitudes, where adults are found close to running water. Adults are predaceous on other small and soft-bodied insects. They are also known to prey on the adults and larvae of biting black flies (Simuliidae). The larvae live in damp soil and among lichens, but some are aquatic. At present, 102 Spilogona species are known from the Palaearctic region (PONT 1986), of which only 51 species are known from Russia. Moreover, most species are recorded from the European territory of Russia. The basic monographs on the Muscidae including the genus Spilogona are by HENNIG (1955­1964) on the entire Palaearctic region (but it is very sketchy outside Europe), GREGOR et al. (2002) on Central Europe, SHINONAGA (2003) on Japan, and XUE (1998) on China. This paper summarises earlier reports of Spilogona species and gives a number of new records and new species from the North European Territory of Russia (Map 1). This territory of Russia is most favourable for Spilogona and is also better studied than the vast area east of the Urals. 29


SOROKINA: The genus Spilogona SCHNABL (Muscidae) in north European Russia

Map 1: Localities of Spilogona SCHNABL in the North European Territory of Russia.

The collections of A. A. STACKELBERG from the St Petersburg district and the collections of V. I. SYCHEVSKAYA from the Murmansk district were for many years the basic collections of Spilogona from the North European Territory of Russia. These collections are kept in the Zoological Museum of the Moscow State University and in the Zoological Museum of the Zoological Institute RAS (St Petersburg). These collections were used for HENNIG's Palaearctic monograph (1955­1964) and for the key by ZIMIN & EL'BERG (1970) to Muscidae from the European part of the former USSR. There is a little further information about Spilogona in this region by R. FREY (1937), R. KROGERUS (1932), C. LUNDSTRæM (1913), and L. TIENSUU (1933, 1936). These papers were based on collections of flies and other insects from the Kanin Peninsula, Kareliya Republic (Sortavala) and St Petersburg district (Ingria). Most recently, a collection of Spilogona was made by Nikita VIKHREV in the Archangel'sk district around Naryan-Mar in the summer of 2008, and this material was kindly entrusted to me for study. Two species new to science were found in VIKHREV's material. Spilogona genualis HUCKETT, Spilogona norvegica (RINGDAHL) and Spilogona nitidicauda (SCHNABL) were found to be new records for the North European Territory of Russia. Spilogona genualis HUCKETT was previously known only from Alaska, Spilogona norvegica (RINGDAHL) was recorded from Eastern Siberia and Spilogona nitidicauda (SCHNABL) was recorded from Western Siberia. 30


Studia dipterologica 16 (2009) Heft 1/2: 29­39

A list is given in the text below of 37 Spilogona species which are currently known from the North European Territory of Russia. This list includes the material studied from the Zoological Museum of the Moscow State University and the Zoological Institute RAS (St Petersburg), the new material from the Archangel'sk district, and species recorded in the literature. The descriptions of two new species are given after the list of species. Materials and methods Spilogona material from the Archangel'sk district is deposited in the Zoological Museum of the Moscow State University (ZMUM) and in the Institute of Systematics and Ecology of Animals, Russian Academy of Sciences, Siberian Branch (Novosibirsk) (SZNM). Other material is also in the Zoological Institute RAS (St Petersburg) (ZIN). Absolute measurements are used for the body length in millimetres (mm). The following abbreviations are used in the text below: acr ­ acrostichal seta, dc ­ dorsocentral seta, a ­ anterior seta, av ­ anteroventral seta, ad ­ anterodorsal seta, p ­ posterior seta, pd ­ posterodorsal seta, pv ­ posteroventral seta, d ­ dorsal seta, v ­ ventral seta. Results List of Spilogona species from the North European Territory of Russia Spilogona acrostichalis (S
Literature: ZIMIN & EL'
BERG

TEIN

, 1916)

(1970): 547.

Spilogona aerea (FALLèN, 1825)
Literature: ZIMIN & EL'
BERG

(1970): 545.

Spilogona almqvistii (HOLMGREN, 1880)
Literature: ZIMIN & EL'
BERG

(1970): 543 [Novaya Zemlya].

Spilogona baltica (R
Literature: H
ENNIG

INGDAHL

, 1918)
(1970): 538.

(1959a): 279 [St Petersburg district]; ZIMIN & EL'

BERG

Material examined: RUSSIA: 2 (ZIN), Leningradskaya oblast', Luzhskiy rayon, 15 km N Luga, Tolmachevo, 18.viii.1935, A. STACKELBERG. 1 (ZIN), vill. Gobzhytsy, 19.viii.1934, A. STACKELBERG.

Spilogona brunneifrons R
Literature: ZIMIN & EL'
BERG

INGDAHL

, 1931

(1970): 543.

Spilogona brunneisquama (ZETTERSTEDT, 1845)
Literature: ZIMIN & EL'
BERG

(1970): 550.

Material examined: RUSSIA: 1 (ZMUM), Murmansk district, Laplandskiy reserve, 10.viii.1975, V. SYCHEVSKAYA.

31


SOROKINA: The genus Spilogona SCHNABL (Muscidae) in north European Russia

Spilogona carbonella (ZETTERSTEDT, 1845)
Literature: ZIMIN & EL'
BERG

(1970): 541.

Spilogona contractifrons (ZETTERSTEDT, 1838)
Literature: LUNDSTRæM & (FREY) 1913: 16 [Kambalnitsa and Ludovoty (Kanin Peninsula, NW USSR), determined by STEIN]; HENNIG (1959b): 289 [Murmansk coast: Alexandrovsk; St Petersburg district]; TIENSUU (1936b): 165 [Sortavala (formerly Finland, now NET)]; ZIMIN & EL'BERG (1970): 538. Material examined: RUSSIA: 2 1 (ZMUM), Murmansk district, Laplandskiy reserve, 22.vii.1975, V. SYCHEVSKAYA. 1 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 30.vi.1931, A. STACKELBERG. 7 (ZIN), Luzhskiy rayon, 15 km N Luga, Tolmachevo, 17.vi.1935, A. STACKELBERG. 1 (ZIN), vill. Bol'shaya Yshchera, 11.viii.1958, A. STACKELBERG. 13 2 (ZMUM, SZNM), Archangel'skaya oblast', Nenetskiy AO, Naryan-Mar, 7­12.vii.2008, . ZEROV, N. VIKHREV. 7 (ZMUM, SZNM), 60 km N Naryan-Mar, Pechora river, 10.vii.2008 and 11.vii.2008, . ZEROV, N. VIKHREV.

Spilogona denigrata (MEIGEN, 1826)
Literature: H
ENNIG

(1959b): 291 [St Petersburg district]; ZIMIN & EL'

BERG

(1970): 538.

Material examined: RUSSIA: 1 (ZMUM), Murmansk district, Laplandskiy reserve, 24.vii.1975, V. SYCHEVSKAYA. 1 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 30.vii.1933, A. STACKELBERG. 3 (ZIN), Luzhskiy rayon, Luga, 4.vii.1932, A. STACKELBERG. 1 (ZIN), vill. Bol'shaya Yshchera, 11.viii.1958, A. STACKELBERG. 1 (ZIN), vill. Gobzhytsy, 5.vi.1934, A. STACKELBERG.

Spilogona depressula (ZETTERSTEDT, 1845)
Literature: H
ENNIG

(1959b): 295 [St Petersburg district]; ZIMIN & EL'

BERG

(1970): 541.

Material examin ed : RU S S IA : 5 (ZIN ), Leningrads kaya oblas t', V s evolozhs kiy rayon, vill. Y kki, 7.vi.1931, 25.vi.1931, 30.vi.1931, 21.viii.1931, A. STACKELBERG. 3 (ZIN), Luzhskiy rayon, vill. Gobzhytsy, 12.viii.1931, A. STACKELBERG. 1 (ZMUM), Archangel'skaya oblast', Nenetskiy AO, Naryan-Mar, 67,6ºN 52,9ºE, 8.vii.2008, . ZEROV.

Spilogona dispar (FALLèN, 1823)
Literature: H
ENNIG

(1959b): 299 [St Petersburg district (as funeralis)]; ZIMIN & EL'

BERG

(1970): 538 (as funeralis).

Material examined: RUSSIA: 12 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 2­11.vii.1931, A. STACKELBERG. 12 1 (ZIN), Luzhskiy rayon, Luga, 22.vii.1955, A. STACKELBERG.

Spilogona falleni PONT, 1984
Literature: H
ENNIG

(1959b): 310 [St Petersburg district (as litorea)]; ZIMIN & EL'

BERG

(1970): 540 (as litorea).

Material examined: RUSSIA: 1 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 2­25.vi.1931, A. STACKELBERG. 1 (ZIN), Luzhskiy rayon, Luga, 5.viii.1934, A. STACKELBERG.

Spilogona genualis HUCKETT, 1965
Material examined: RUSSIA: 5 1 (ZMUM, SZNM), Archangel'skaya oblast', Nenetskiy AO, Naryan-Mar, 67,6ºN 52,9ºE, 8.vii.2008, . ZEROV, N. VIKHREV.

Spilogona karelica (TIENSUU, 1936)
Literature: TIENSUU (1936a): 34 [Sortavala (formerly Finland, now NET)]; F EL'BERG (1970): 538.
REY

(1937): 251 [Sortavala]; ZIMIN &

32


Studia dipterologica 16 (2009) Heft 1/2: 29­39

Spilogona marginifera HENNIG, 1959
Literature: H
ENNIG

(1959b): 314 [St Petersburg district]; ZIMIN & EL'

BERG

(1970): 538.

Material examined: RUSSIA: 1 3 (ZIN), Leningradskaya oblast', Luzhskiy rayon, Luga, 12.vii.1935, 23.vii.1953, A. STACKELBERG; 1 (ZIN), Vsevolozhskiy rayon, vill. Ozerki, 27.vii.1919, A. STACKELBERG.

Spilogona megastoma (BOHEMAN, 1866)
Literature: ZIMIN & EL'
BERG

(1970): 543 [Novaya Zemlya].

Spilogona nigriventris (ZETTERSTEDT, 1845)
Literature: H
ENNIG

(1959b): 319 [Murmansk coast: Alexandrovsk]; ZIMIN & EL'

BERG

(1970): 545.

Spilogona nitidicauda (S

CHNABL

, 1911)

Material examined: RUSSIA: 1 (ZMUM), Archangel'skaya oblast', Nenetskiy AO, 60 km N Naryan-Mar, Pechora river, 68,3ºN 53,3ºE, 11.vii.2008, . ZEROV.

Spilogona nordenskioldi (HOLMGREN, 1883)
Literature: ZIMIN & EL'
BERG

(1970): 545 [Novaya Zemlya].

Spilogona norvegica (R

INGDAHL

, 1932)

Material examined: RUSSIA: 1 (ZMUM), Archangel'skaya oblast', Nenetskiy AO, Naryan-Mar, 67,6ºN 52,9ºE, 6.vii.2008, . ZEROV. 3 (ZMUM, SZNM), same locality data, 7.vii.2008, N. VIKHREV.

Spilogona novemmaculata (ZETTERSTEDT, 1860)
Literature: ZIMIN & EL'
BERG

(1970): 546.

Spilogona obscuripennis (S
Literature: ZIMIN & EL'
BERG

TEIN

, 1916)

(1970): 545.

Spilogona pacifica (MEIGEN, 1826)
Literature: TIENSUU (1933): 122 [Sortavala (formerly Finland, now NET; as nupta)]; HENNIG (1959c): 357 [St Petersburg district (as vana)]; ZIMIN & EL'BERG (1970): 550 (as vana). Material examined: RUSSIA: 5 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 18.vi.1931, 2.vii.1931, 15.vii.1931, 27.vii.1931, 9.vii.1932, A. STACKELBERG. 2 (ZIN), Luzhskiy rayon, Luga, vill. Gobzhytsa, 4.vi.1934, A. STACKELBERG.

Spilogona pseudodispar (FREY, 1915)
Literature: ZIMIN & EL'
BERG

(1970): 544.

Spilogona quinquelineata (ZETTERSTEDT, 1838)
Literature: H
ENNIG

(1959b): 330 [St Petersburg district]; ZIMIN & EL'

BERG

(1970): 549.

Material examined: RUSSIA: 4 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 7­16.vii.1933, A. STACKELBERG.

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SOROKINA: The genus Spilogona SCHNABL (Muscidae) in north European Russia

Spilogona scutulata (S

CHNABL

, 1911)
BERG

Literature: FREY (1937): 251 [Sortavala (formerly Finland, now NET; as munda)]; ZIMIN & EL'

(1970): 550.

Spilogona semiglobosa (R

INGDAHL

, 1916)
BERG

Literature: TIENSUU (1933): 122 [Sortavala (formerly Finland, now NET)]; ZIMIN & EL'

(1970): 538.

Spilogona septemnotata (ZETTERSTEDT, 1845)
Literature: ZIMIN & EL'
BERG

(1970): 547.

Spilogona sjostedti (R
Literature: ZIMIN & EL'
BERG

INGDAHL

, 1926)

(1970): 545.

Material examined: RUSSIA: 1 (ZMUM), Archangel'skaya oblast', Nenetskiy AO, 60 km N Naryan-Mar, Pechora river, 68,3ºN 53,3ºE, 11.vii.2008, . ZEROV.

Spilogona sordidipennis (HOLMGREN, 1883)
Literature: ZIMIN & EL'
BERG

(1970): 542 [Novaya Zemlya].

Spilogona surda (ZETTERSTEDT, 1845)
Literature: TIENSUU (1936b): 165 [Sortavala (formerly Finland, now NET)]; H district]; ZIMIN & EL'BERG (1970): 538.
ENNIG

(1959c): 347 [St Petersburg

Material examined: RUSSIA: 2 (ZIN), Leningradskaya oblast', Vsevolozhskiy rayon, vill. Ykki, 14.vi.1932 and 18.vi.1932, A. STACKELBERG.

Spilogona tenuis HENNIG, 1959
Literature: H
ENNIG

(1959c): 348 [paratypes from St Petersburg district]; ZIMIN & EL'

BERG

(1970): 550.

Material examined: RUSSIA: 1 (paratype in ZIN), Leningradskaya oblast', Luzhskiy rayon, 15 km N Luga, Tolmachevo, 22.vi.1937, A. STACKELBERG.

Spilogona triangulifera (ZETTERSTEDT, 1838)
Literature: LUNDSTRæM & FREY (1913): 16 [Kanin peninsula (NW USSR), determined by S (1970): 542 [Novaya Zemlya].
TEIN];

ZIMIN & EL'

BERG

Material examined: RUSSIA: 1 (ZMUM), Archangel'skaya oblast', Nenetskiy AO, 60 km N Naryan-Mar, Pechora river, 9.vii.2008, N. VIKHREV. 3 (ZMUM, SZNM), Kashin island, 68,2ºN 53,8ºE, 10.vii.2008, . ZEROV.

Spilogona trigonata (ZETTERSTEDT, 1838)
Literature: ZIMIN & EL'
BERG

(1970): 545.

Spilogona tundrae (S
Literature: H
ENNIG

CHNABL
BERG

, 1915)

(1959c): 355 [Kanin Peninsula]; ZIMIN & EL'

(1970): 543.

Material examined: RUSSIA: 1 (ZMUM), Archangel'skaya oblast', Nenetskiy AO, 60 km N Naryan-Mar, Pechora river, 11.vii.2008, N. VIKHREV.

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Studia dipterologica 16 (2009) Heft 1/2: 29­39

Spilogona varsaviensis (S
Literature: K
ROGERUS

CHNABL
BERG

, 1911)

(1932): 138 [Dunes of Ingria]; ZIMIN & EL'

(1970): 538.

Spilogona veterrima (ZETTERSTEDT, 1845)
Literature: K
ROGERUS

(1932): 138 [probably present in the dunes of Ingria]; ZIMIN & EL'

BERG

(1970): 550.

Spilogona vikhrevi spec. nov.
(Figs 1­3) Material examined: Holotype: , RUSSIA: Archangel'skaya oblast', Nenetskiy AO, 60 km N Naryan-Mar, Pechora river, 10.vii.2008, N. VIKHREV (ZMUM). Paratype: 1 , same data as holotype (SZNM).

Description. Length: body 6.0­6.3 mm, wing 5.2­5.5 mm. Male. Head: Ground-colour black. Frons, fronto-orbital plate, parafacial and gena brownishyellow pruinose. Mouth-edge not projecting beyond level of profrons. Eye bare. Fronto-orbital plates separated by a narrow frontal vitta. Frontal vitta narrower than diameter of anterior ocellus. Distance between eye margins at upper part of frons about as wide as the distance between posterior ocelli. Frons with 9­11 pairs of frontal seta (only the first pair stronger than the others). Width of parafacial at level of base of antenna as wide as postpedicel, at middle 2/3 of width

2

3

Figs 1 ­3: Spilogona vikhrevi spec. nov. ­ 1: Sternite 5; ­ 2: Genitalia, lateral view; ­ 3: Cercal plate and surstylus, posterior view. Scale bars = 0.25 mm.

1

35


SOROKINA: The genus Spilogona SCHNABL (Muscidae) in north European Russia

of postpedicel. Height of gena 2 times width of postpedicel. Antenna black, postpedicel ca. 1.3 times as long as wide. Arista black, short pubescent, with longest individual hairs much shorter than basal diameter of arista. Palpus black. Proboscis short, with two long and strong spines, prementum completely dusted. Labella small. Thorax: Ground-colour black. Scutum, scutellum and pleura brownish-yellow dusted. Groundsetulae of scutum relatively long and dense, all setae hair-like, difficult to differentiate from the ground-setulae. Presutural acr in 2 rows, 4(5)+5(6) dc (two pairs of setae near scutellum longer and stronger than the others). Notopleuron bare. Anepisternum with one long and strong interspatial seta. Katepisternal setae 1+1. Scutellum without preapical hairs on upper border of declivities. Wing dark with blackish base, all veins more or less infuscated. Calypters dark: upper one black, lower one brown. Haltere yellow. Legs: Ground-colour black. Fore tibia without p, anterior surface with numerous short strong setulae. Mid femur with 6 pv on basal half; 0 a and 2 pd-p preapical setae. Mid tibia with 1 ad, 2 pd, 2 av, 1 pv, 1 short v. Hind femur with 7 av, 9 ad, 5 short pv on basal half, 2 pd at apex. Hind tibia with 2 av, 1 ad, 1­2 short pd, 0 pv; 1 ad apical, half as long as d apical, 1 av apical which is 2/3 as long as d apical. Abdomen: Black with brownish-yellow dust, elongated, cylindrical. Tergites 3­5 with indistinct diffuse marks which are a little darker than abdominal colour. Sternite 1 bare (one male has one hair on sternite 1), sternite 5 as in figure 1. Terminalia: as in figures 2, 3. Female. Unknown. Distribution. Russia, Archangel'sk district. Etymology. The species name is a patronym in honour of the dipterist Nikita VIKHREV (Moscow), who collected the type series. Diagnosis. The species is very similar to Spilogona tundrica (SCHNABL). The male of the new species can be distinguished as follows: hind femur with short pv, black upper calypter and brown lower calypter, yellow haltere, postpedicel ca. 1.3 times as long as wide, dusted prementum, dark abdomen with indistinct diffuse marks, hair-like acr and fine dc, one long anepisternal interspatial seta. The new species can be incorporated into HENNIG's key to males of Palaearctic Spilogona (HENNIG 1959: 262) by the following couplets: 144 (143) as in HENNIG 145a (145b) Calypters blackish ........................................................... S. vikhrevi spec. nov. 145b (145a) Calypters yellow or white ............................................................................ 145 145 (146) as in HENNIG If the pv setae on hind femur are considered to be indistinct, then the new species can be incorporated into HENNIG's key (HENNIG 1959: 263) by the following couplets: 177 (188) as in HENNIG 177a (178) Calypters blackish or brownish .................................................................. 179a 178 (177a) Calypters yellow or white ............................................................................ 180 179a (179) Haltere brown .................................................................... S. tundrica SCHNABL 179 (179a) Haltere yellow ................................................................. S. vikhrevi spec. nov. 180 (181) as in HENNIG

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Studia dipterologica 16 (2009) Heft 1/2: 29­39

6 5

Figs 4­6: Spilogona archangelica spec. nov. ­ 4: Sternite 5; ­ 5: Genitalia, lateral view; ­ 6: Cercal plate and surstylus, posterior view. Scale bars = 0.25 mm.

4

Spilogona archangelica spec. nov.
(Figs 4­6) Material examined: Holotype: , RUSSIA: Archangel'skaya oblast', Nenetskiy AO, around Naryan-Mar, 8.vii.2008, N. VIKHREV (ZMUM). Paratype: 1 , same data as holotype (SZNM).

Description. Length: body 5.0­5.2 mm, wing 4.6 mm. Male. Head: Ground-colour black. Fronto-orbital plate and parafacial densely silvery pruinose, gena yellowish-grey or silvery pruinose. Mouth-edge not projecting beyond level of profrons. Eye bare. Fronto-orbital plates touching in middle; fronto-orbital plate as wide as or wider than diameter of anterior ocellus. Distance between eye margins at upper part of frons about as wide as the distance between posterior ocelli. Frons with 7­8 pairs of frontal seta (last pair half as long as the others). Width of parafacial at level of base of antenna as wide as postpedicel, at middle about 1/2 of width of postpedicel. Height of gena about 2 times width of postpedicel. Antenna black, postpedicel ca. 2 times as long as wide. Arista black, pubescent, with longest individual hairs as long as or a little longer than basal diameter of arista. Palpus black. Proboscis short, prementum shining. Labella large. Thorax: Ground-colour black. Scutum, scutellum and pleura more or less densely grey dusted. Scutum without distinct longitudinal stripes. Presutural acr in 2 uneven rows, 2+3(4) dc. Notopleuron bare. Anepisternum without interspatial seta. Katepisternal setae 1+2 (lower seta 37


SOROKINA: The genus Spilogona SCHNABL (Muscidae) in north European Russia

weak). Scutellum without preapical hairs on upper border of declivities. Wing light, with a slight brownish tinge, not darked at base. Costa with weak spinules. Calypters yellow. Haltere yellow. Legs: Ground-colour black. Knees brownish. Fore tibia with 1 strong p. Mid femur with 4­6 pv on basal half; without or with 1 weak a and 3 strong pd-p preapical setae. Mid tibia with 1 ad, 2 pd, without any v. Hind femur with a complete row of ad, 3­4 av on apical half, 8 long hair-like p on basal half, without pv. Hind tibia with 1­2 av, 1­2 ad, without pd or pv; 1 ad apical, half as long as d apical. Abdomen: Black with relatively thick yellowish-grey dust, cylindrical. Tergites 3 and 4 with small brown diffuse lateral marks, more or less triangular. Sternite 1 bare, sternite 5 as in figure 4. Terminalia: as in figures 5, 6. Female. Unknown. Distribution. Russia, Archangel'sk district. Etymology. The name `archangelica' is an adjective derived from the medieval Latin word `angelicus' with the prefix `arch-'. This reflects the name of the type locality, Archangel'sk, which was originally a small settlement named after a monastery at the mouth of the River Dvina dedicated to the Archangel Michael. Diagnosis. The species is very similar to Spilogona kuntzei (SCHNABL) and Spilogona varsaviensis (SCHNABL). The male of the new species can be distinguished as follows: width of fronto-orbital plate as wide as or wider than diameter of anterior ocellus, abdomen yellowish-grey dusted, with diffuse lateral brownish marks, fore tibia with 1 strong p, mid tibia with 1 strong ad, hind tibia without pd, prementum shining, haltere yellow, katepisternal setae 1+2. If the new species has 3 postsutural dc, then it can be incorporated into HENNIG's key to males of Palaearctic Spilogona (HENNIG 1959: 262) by the following couplets: 54 (53) as in HENNIG 55 (56) Width of parafacial at middle about as 1/2 of width of postpedicel ............. 55a 55a (55b) Fore tibia without p; mid tibia without ad; abdomen with distinct dark marks ... ...................................................................................... S. varsaviensis SCHNABL 55b (55a) Fore tibia with a distinct p; mid tibia with a strong ad; abdomen with diffuse lateral marks ............................................................ S. archangelica spec. nov. 56 (55) as in HENNIG If the new species has 4 postsutural dc, then it can be incorporated into HENNIG's key to males of Palaearctic Spilogona (HENNIG 1959: 262) by the following couplets: 214 (215) as in HENNIG 215 (215a) Hind femur with elongated pv .................................................................... 215a 215a (215b) Fore tibia without p; mid tibia without or with a short ad; abdomen whitish-blue dusted with large trapezoid marks ....................................... S. kuntzei SCHNABL 215b (215a) Fore tibia with a distinct p; mid tibia with a strong ad; abdomen yellowish-grey dusted with small triangular marks ......................... S. archangelica spec. nov. 216 (213) as in HENNIG Acknowledgements
I am grateful to Nikita VIKHREV (Moscow, Russia) for the opportunity of studying the material from the Archangel'sk district. I would also like to thank Dr A. L. OZEROV (Moscow, Russia) and Dr E. P. NARCHUK (St Petersburg, Russia)

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Studia dipterologica 16 (2009) Heft 1/2: 29­39 for the opportunity of studying the collections of the Zoological Museum of the Moscow State University and the Zoological Institute RAS (St Petersburg). I would especially like to thank Dr A. C. PONT (Oxford, UK) for discussion of this manuscript and for giving me access to his personal notes on Spilogona, and also for correcting the English of this paper.

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REY,

Author's address Vera S. SOROKINA Zoological Museum Institute of Systematics and Ecology of Animals Russian Academy of Sciences, Siberian Branch Frunze Street 11 Novosibirsk, 630091 Russia E-mail: sorokinavs@mail.ru
The paper was accepted on 11 May 2010. Editum: 29 December 2010.

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