Äîêóìåíò âçÿò èç êýøà ïîèñêîâîé ìàøèíû. Àäðåñ îðèãèíàëüíîãî äîêóìåíòà : http://www.sevin.ru/dissertations/ecology/51.pdf Äàòà èçìåíåíèÿ: Fri Nov 6 13:53:14 2009 Äàòà èíäåêñèðîâàíèÿ: Tue Oct 2 11:10:25 2012 Êîäèðîâêà:

(INSECTA, COLEOPTERA)

03.00.09 ­



2009


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«8 » 2009 . 14 002.213.01 . .. : 119071, , , 33 (495) 952-35-84 e-mail: admin@sevin.ru : 119071, , , 33 « » 2009 .

, ..


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. Arthropoda , . «» , .. (Bourdeaux, 1979; Bitsch, 2002). , , , Crustacea Myriapoda, Hexapoda (Apterygota). Pterygota, Coleoptera , , , . , (Joshi, 1976), (Crowson, 1938, 1944). Coleoptera, (Baehr, 1975, 1979; Beutel, 1992; Friedrich et al, 2006, 2009). . . - Coleoptera . : 1) , ; ; ; 2) , , ; 3) ;


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4) , - Coleoptera; 5) Coleoptera ; 6) Coleoptera. . 202 90 , . , . . Coleoptera , Neuropteroidea. Coleoptera Holometabola . . . . Coleoptera. , , Coleoptera. - , . . : XIII (, 2007); XIII European Carabidologists Meeting, (Blagoevgrad, 2007),


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- . . 6 , 3 , 1 ­ ; 1 2 ­ . . , 7 , . 189 , 4 196 . 160 , 145 . . .. () . .. () , , , .. (), .. (), .. (), .. (), .. (), .. (), .. (), .. (), Killian A. (Wroclaw University, ), Pollock D.A. (Eastern New Mexico University, ), lipiski S.A. (CSIRO Entomology, ), Horak J. (), (-): .., .., .. .. .


. , . 1. 1.1. Coleoptera

, , , , , XIX (Eschscholtz, 1820; Baer, 1826; Audouin, 1828; Mac Leay, 1830; Faivre, 1861), (Baehr, 1979; Ge et al, 2007). . (Berlese, 1909),


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. . (Crampton, 1918), Pterygota. , , . , Coleoptera, - , (Speyer, 1922; Crome, 1957; Marvillet, 1960; Jaboulet, 1960; Barlet, 1992). , , (Paulian, 1944), 14 . , , Coleoptera. , , . , , , Coleoptera , . 1.2. Coleoptera Coleoptera, (Chapius, CandÈze, 1853; Perris, 1877; SchiÆdte, 1862-1883; Saalas, 1917, 1923; Verhoeff, 1923). , , , , .. , , (BÆving, Craighead, 1931).


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Coleoptera , . (Crowson, 1955; Rozen, 1960; Viedma, 1966; Doyen, Lawrence, 1979; Young 1976, 1985; Lawrence & lipinski, 1999). ( ) (Bousquet, 1984; Kovarik & Passoa, 1993; Makarov, 1996; Delgado, 1996; Solodovnikov, 2007) (Ruhnau, 1984; Beutel et al, 1993, 1998, 2000; Kiselyova, McHugh, 2006; Zaitsev, 2005, 2008). , Coleopera . , , . , , , (van Emden, 1957; Hinton, 1955, 1966). Coleoptera, , 80- XX . , Hydradephaga (Ruhnau, 1984). . (Beutel, 1988, 1991; Beutel et al, 1999a, 1999b, 2000, 2006) Coleoptera, . ; , , , - . Coleoptera . , (Crowson, 1938) .


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2. 202 90 Coleoptera Archostemata, Adephaga Polyphaga, 70% . Ascalaphus sp. (Neuroptera, Ascalaphidae), Myrmeleon sp. (Neuroptera, Myrmeleontidae), Osmylus sp. (Neuroptera, Osmylidae), Chrysopa sp. (Neuroptera, Chrysopidae), Inocellia sp. (Rhaphidioptera, Inocelliidae), Sialis sp. (Megaloptera, Sialidae), Panorpa communis L. (Mecoptera, Panorpidae), Tenthrediniidae gen. sp. (Hymenoptera), Allotrichia sp. (Trichoptera, Hydroptilidae). , . , NaOH . . Leica MZ6.5 MZ16.5, . Corel Draw 12. Statistica 8.0. , U- - ( - ) 95% (, 2001). 3. Coleoptera 3.1. Coleoptera - , . Coleoptera 10 , . (. 1).


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. (aes1) . (aem1), . Coleoptera , . (apl1), . . , . (f1) - , , . ( . 1 ), - . (ais1), - , , . - . , , ( , ), .


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, , , . , , ( ) . (spina), - - , , . ( , ) , . . 3.2. Coleoptera 3.2.1. ( Calosoma auropunctatum Herbst) Carabidae . , , Calosoma auropunctatum Herbst . , Calosoma , , . , , - - . 3.2.2. ( Cerambycidae) , , (Buprestidae, Curculionidae, Cerambycidae). , , , . . , .


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, (Oedeomeridae, Melandryidae, Peltidae) ­ , , . , , . 3.3. 3.3.1. 3-5 , . Coleoptera, , : , , . . Calosoma auropunctatum Herbst., Ophonus asureus Fabr., Carabus granulatus L., Anisodactylus binotatus Fabr., Platynus assimilis Payk. (Carabidae) Cucujus haematodes Er. (Cucujidae). , apl1, aes2-3 f1-3 (2-3) . , , ais1-3 , , -, . , , , . : Carabus nemoralis Muller, C. granulatus L. (Carabidae), Stenagostus undulatus Deg. (Elateridae), Dorcadion sp. (Cerambycidae), Hylecoetus dermestoides L. (Lymexylidae), Schizotus pectinicornis L. (Pyrochroidae), Calopus serraticornis L. (Oedeomeridae). . ,


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Stenagostus Carabus ( ) . , , ( ) , de novo ( , , Stenagostus) (Carabus). f3 . (.. , ). , , : f1-2, f3 ( ), . -, , . , , ( ). 3.3.2. Holometabola Coleoptera Holometabola. (Poyarkoff, 1914; Hinton, 1955; Sehnal et al, 1996). , () , , , . , Holometabola, Coleoptera. , , «» Holometabola. , , , (Lubbock, 1883; Berlese, 1913; , 1929; Henson, 1946, , 1991).


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Hemi- Holometabola (. 2). «» , , Coleoptera . , , , (Poyarkoff, 1914; Henson, 1946). , , . ( ), . , « , » (, 1961) , .. Hemimetabola.

4. Neuropteroidea, Mecoptera, Hymenoptera, Lepidoptera Trichoptera Neuropteroidea (Neuroptera, Rhaphidioptera Megaloptera) Coleoptera ( ), . . , Sialis Coleoptera. : Hymenoptera Lepidoptera, Mecoptera, , Trichoptera. Panorpa Allotrichia ; -


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, ­ aes2-3 sp3 . , Mecoptera Trichoptera Neuropteroidea: , . , Coleoptera Neuropteroidea, Megaloptera, (Crampton, 1914; Beutel, 2007). 5.

. , , . , .

6. Coleoptera 6.1. Coleoptera , Coleoptera : 1) , 2) . , , . , Schizotus, Dendroides, Pyrochroa Pyrochroidae, , ; Melandrya Orchesia (Melandryidae) . , Coleoptera, , , , . , Tenebrionoidea Cucujoidea, .


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. , , , . , , ( ­ ) . 6.2. Coleoptera , , . 8 , , , ( 1). . , ; ­ . . , , , . , «». ( 19 ) , , apl1 aes1-3. , , ( ) , . , , - , . , - Silphidae, Eulichadidae, Endomychidae - Scarabaeoidae. ,


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Trachypachydae, Carabidae, Hydrophilidae, Erotylidae

apl1

aes1-3 f2-3 apl2-3 sp2-3

ais2-3

ais1




Silphidae, Eulichadidae, Endomychidae, Scarabaeoidae, Endecatomidae Lampyridae Coccinellidae Nosodendridae Dytiscidae, Gyrinidae, Lucanidae Byrrhidae, Dascillidae, Scirtidae, Psephenidae, Helophoridae, Passalidae, etc Lymexylonidae, Cerophytidae Staphylinidae, Scaphidiidae Heteroceridae, Elmidae Cantharidae, Callirhipidae Elateridae, Drilidae, Lycidae, Ptinidae, Dermestidae, Haliplidae










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Zopheridae, Oedemeridae, Pythidae, Monotomidae, Monommatidae, Othnidae Salpingidae, Synchroidae, Nitidulidae, Leiodidae, Corylophidae, Pyrochroinae Cryptocephalinae Cleridae, Peltidae, Trogossitidae, Silvanidae Noteridae, Rhysodidae, Anobiidae, Malachidae, Cucujidae, Byturidae, Ciidae, etc

















1. Coleoptera. . ­ ; . ,


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. , , . 6.3. Coleoptera , , , ( ) . , , , . , , , - . Coleoptera ( 2). , , , , . , , , . -, . -, ­ , / . -, , .


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Lawrence, Newton, 1995 \. Carabidae (Bell, 1998); Carabidae (Beutel, 1990, 1992); Rhysodes Rhysodidae, Adephaga ( Carabidae) Adephaga (Ponomarenko, 1995); Archostemata (Makarov, 2008) Nodinus Drapetes Orphilus Apateticinae, Staphylinidae Lissominae, Elateridae Orphilinae, Dermestidae Apateticinae, Silphidae (Madge, 1980) Lissomidae (Burakowski, 1976 ) Nosodendridae (Ivie, 1985, 2002) Apateticinae, Staphylinidae, Scaphidiidae Lissominae, Elateridae Orphilinae, Dermestidae; Thylodriinae Carabidae

Lagria

Lagriinae, Tenebrionidae

Lagriidae (Boving, 1931; Crowson, 1981)

Lagriinae, Tenebrionidae

Elacatis Inopeplus, Diagrypnodes Aglenus Pedilus

Othniinae, Salpingidae Inopeplinae, Salpingidae Agleninae, Salpingidae Pedilinae, Pyrochroidae

Othniidae (Boving, 1931); Elacatidae (Crowson, 1981) Inopeplidae (Sato, Hatta, 1988) Agleninae, Colydiidae (Dajoz, 1977) Anthicidae (Abdullah, 1969); Pedilidae (Lacordaire, 1859) Pythidae (Watt, 1987);

Othniinae, Pythidae ( ) Inopeplidae Agleninae, Salpingidae Pedilinae, Pyrochroidae

Morpholycus, Temnopalpus

Pilipalpinae, Pyrochroidae Pilipalpidae (Nikitsky, 1986)

Pilipalpidae

Colydiidae, Zopheridae, Monommatidae Meryx, Ulodes

Colydiidae, Zopheridae, Monommatidae

\. Zopheridae (Lawrence, lipinski, 1997)

Zopheridae (Colydiinae + Zopherinae), Monommatidae

Ulodidae

Ulodinae, Zopheridae (Doyen, Lawrence, 1979)

Ulodidae, Archeocrypticidae Mycetophagidae



2. Coleoptera. (Lawrence et al, 1995) ; .


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7. Coleoptera Coleoptera . , , . , (, 1974, 1976). () () . «» . , , Adephaga, Staphylinoidea Hydrophyloidea, , . ­ , . , , - Buprestidae, Cerambycidae, Curculionidae, . , , , . , , . Coleoptera. , , . Megaloptera (Crampton 1918; Wheeler et al, 2001, , 2006), , Coleoptera . ­ Archostemata , , , (Beutel, HÆrnschemeyer, 2008).


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. , , , , . Adephaga ( Trachypachidae Gyrinidae) Hydrophyloidea Polyphaga (, 1969, 1995; Grimaldi, Engel, 2005). , Trachypachidae (Geadephaga), Dytiscidae, Gyrinidae (Hydradephaga) Hydrophilidae Polyphaga . 7.1. Coleoptera

. , , , . Hydrophiloidea Elateroidea. , ­ . ( ) Coleoptera , -, - , . Polyphaga ( Cleroidea, Drilidae, Elateridae Tenebrionidae) , . , Adephaga. , , . - , . , , Buprestidae, Cerambycidae Curculionidae. Coleoptera


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. , , . , , . «», (, 1996). , . . , , , . , , Cicindelini Carabidae, Clytrini Chrysomelidae, Callirhipidae, Eulichadidae . , , . , Coleoptera ­ , . , (, 1929, 1939). , . . , . , . , , , , , , . , (, 1939; , 1991), Holometabola .


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1. Coleoptera , . 2. , . . , . 3. Coleoptera , , . , Coleoptera , -, Holometabola, , , Hemimetabola 4. 8 , , ­ 5. / . . , 6. Coleoptera, Nodynus








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(Staphylinidae, Apateticinae); Lagria (Tenebrionidae, Lagriinae); Inopeplus, Diagrypnodes (Inopeplidae); Elacatis (Pythidae, Othniinae); Pedilus (Pyrochroidae, Pedilinae); Morpholycus, Temnopalpus (Pilipalpidae) . 7. , -, Carabidae, Trachypachidae, Dytiscidae, Gyrinidae (Adephaga), Hydrophilidae (Polyphaga). Sialis, Coleoptera Megaloptera. 8. ­ , . . , , . .. - (Coleoptera, Pyrochroidae) / .. // ­ 2005. - 14 (1). ­ . 15-20. Zaitsev A.A. The endoskeleton of thoracic segments of Carabid larvae (Coleoptera, Carabidae) / .. // Back to the Roots and Back to the Future? Towards a New Synthesis amongst Taxonomic, Ecological and Biogeographical Approaches in Carabidology. - 2007. - Abstract Volume - p. 68. .. Coleoptera / .. // XIII , . ­ 2007. ­ . 117-118. Zaitsev A.A. Description of larval and imaginal stages of new species from the genera Pseudacroleptus Pic, 1911 Ceratoprion Gorham, 1880 (Coleoptera: Lycidae: Leptolycinae) / S.V. Kazantsev, A.A. Zaitsev // Russian Entomol. J. ­ 2008. - 17 (3). ­ . 283-292. Zaitsev A.A. Thoracic endoskeleton of carabid larvae (Coleoptera, Carabidae) / A.A. Zaitsev // Back to the Roots and Back to the Future? Towards a New Synthesis amongst Taxonomic, Ecological and Biogeographical Approaches in Carabidology Proceedings of the XIII


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European Carabidologists Meeting, Blagoevgrad. ­ 2008. ­ . 159-171. .. Tenebrionoidea (Insecta, Coleoptera) / .. // ­ 2009. ­ 8 (2). - . 189-204.