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Russian Entomol. J. 22(1): 7576

ї RUSSIAN ENTOMOLOGICAL JOURNAL, 2013

Description of new species of Thricops Rondani, 1856 (Diptera: Muscidae) from the Altai Mountains Описание нового вида Thricops Rondani, 1856 (Diptera: Muscidae) из горного Алтая Nikita E. Vikhrev Н.Е. Вихрев
Zoological Museum of Moscow University, Bolshaya Nikitskaya 6, Moscow 125009, Russia. E-mail: nikita6510@ya.ru Зоологический музей МГУ им. М.В. Ломоносова, Большая Никитская ул., 6, Москва 125009, Россия.

KEY WORDS: Thricops kosterini, Muscidae, Diptera, new species. КЛЮЧЕВЫЕ СЛОВА: Thricops kosterini, Muscidae, Diptera, новый вид. ABSTRACT. Thricops kosterini sp.n. is described from the Altai Mountains, E Kazakhstan province of Kazakhstan. РЕЗЮМЕ. Thricops kosterini sp.n. описан из горного Алтая, с территории Восточно-Казахстанской области Казахстана. anterior, posterior, dorsal, ventral seta(e); prst presutural, post postsutural. The abbreviation for the tarsi as tar followed by a pair of digits separated by a hyphen was proposed by Vikhrev [2011]: the first digit (1 to 3) gives the leg number and the second digit (1 to 5) the number of the tarsal segment. For example, tar14 4th segment of fore tarsus; tar31 hind basitarsus. Thricops kosterini sp.n. Figs 13.
MATERIAL. Holotype , Kazakhstan: E Kazakhstan reg., Ust-Chindagatuy env., 1750 m asl, 49.25њN 87.00њE, 35.VII.2012, O.Kosterin. Paratype, 1, the same data.

Introduction
Genus Thricops with 45 valid species, is one of the best known and the less problematic genera in Muscidae, partly due to its exclusively Holarctic distribution and partly due to existence of the fresh revision of the world fauna by Savage [2003]. The Russian fauna of Thricops was reviewed by Vikhrev & Sorokina [2009], this paper showed that the genus is rather evenly distributed over the vast territory of Russia except the Caucasus which is the hot spot of the diversity and endemism for Thricops. Recently Sorokina [2012] published her report on Muscidae fauna of the Altai Mountains based on collecting during 6 field seasons. Thus, finding of a new species of Thricops in the Altai Mts. was a surprise, it was collected in the territory of Kazakhstan, but 23 km from the Russian border, 1015 km from the Chinese border and about in 50 km from the area where Sorokina and her colleagues worked in the Russian territory.

Material and methods
The holotype and paratype are in the Zoological Museum of Moscow University (ZMUM). Geographical coordinates are given in the Decimal Degrees format. The following generally accepted abbreviations for morphological structures are used: f1, t1, f2, t2, f3, t3 fore-, mid-, hind- femur or tibia respectively; ac acrostichal setae; dc dorsocentral setae; a, p, d, v

DESCRIPTION. Male, body length 5 mm (Fig. 1). Head holoptic, the narrowest distance between eyes 1.5x diameter of the anterior ocellus. Fronto-orbital plates narrow, grey; interfrontalia linear at middle, black; parafacials half as wide as width of antenna, grey; face grey; gena about 0.2x eye height, grey; occiput grey dusted. Fronto-orbital plates with 7 inclinate setae and some hairs in upper third. Antenna black, short; arista virtually bare. Palpi black; mentum of proboscis thinly dusted. Thorax black with thin grey dusting; scutum in posterior view with black median vitta along ac hairs. Thoracic setae: ac 0+1, prst ac hairs in 4 rows; dc 2+4; postpronotal 2, intraalars 1+2, supraalars 1+1; prealar seta indistinct; notopleuron bare except 2 strong setae; katepimeron bare; scutellum bare ventrally. Wings hyaline, veins bare, halters whitish-yellow; calypters whitish. Legs: tibiae and apices of femora yellow; the rest of femora and tarsi black. t1 without setae except preapicals. tar14 with 23 rather long d setae, otherwise without modifications. f2 in basal 2/3 on av to pv surface with fine setae subequal to femur width. t2 with 2 p setae. Hind coxa with setae on inner posterior margin. f3 with a remarkable modification: a series of 3 strong hooklike pv spines at middle (Fig. 3); a row of p setae with length subequal to femur width, in basal half; 4 strong av seta in apical 1/3. t3 with only single seta a long pd below middle; apex of t3 with 2 modified, stub-like apical pv spines, which are truncated closely to base, as if broken, that is not in the fact case since these stubs are the same on the right and left hind tibiae (normal, not truncated pv spines at apex of t3 present in several species of Thricops).


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Figs 13. Thricops kosterini sp. n., holotype: 1 general view; 2 apex of t3; 3 pv spines on f3. Рис. 13. Thricops kosterini sp. n., голотип: 1 общий вид; 2 вершина t3; 3 pv шипы на f3.

Abdomen densely grey dusted, with black median vitta on tergites 3 and 4. Structure of male genitalia in Thricops has a limited value, there are species with long surstyli and narrow cercal plate, other species have short wider surstyli and wide cercal plate. I had not dissected abdomen of single available male specimen, from outward examination I suppose that T. kosterini sp.n. has terminalia of long and narrow type. Female paratype has both mid legs missing, female differs from male as follows: frons wide, densely whitish-grey dusted, interfrontals absent; tar14 without elongated d setae; thorax more densely grey dusted; short (3 times shorter than posterior notopleural) but distinct prealar seta present; f3 without pv spines and without p setae in basal half; t3 with pd seta shorter than in male but with 1 ad and 2 av setae below middle; abdomen with median vitta indistinct. DIAGNOSIS. Male is unmistakable due to 3 submedian pv spines on f3; the single seta (pd) on t3; pair of ventral stubs of setae at the apex of t3. Female has bicolourous legs: dark femora except of apex, yellow apex of femora and tibiae (the same legs colour is present only in northern specimens of T. genarum (Zetterstedt, 1838). ETYMOLOGY. The species named after its collector Oleg Kosterin (Novosibirsk). RELATIONSHIP. Probably T. kosterini sp.n. is related to T. foveolatus (Zetterstedt, 1845), both species sharing a small size, narrow gena, reduced ac and prealar seta, a simple (for Thricops) leg chaetotaxy. T. foveolatus is rare in collec-

tions, according to my observations in the Caucasus T. foveolatus it is probably associated with horse dung and I especially asked Oleg to try to collect from/near horse dung on the Altai, but it is unknown where and how exactly the new species was found. ACKNOWLEDGEMENTS. I am grateful to people who made the specimens of the new species collected and available: to Oleg Kosterin who collected it, to Aliya Gabdullina, an entomologist from Katon-Karagay, who arraned the trip to the easternmost corner of Kazakhstan and to Nurlan Katkenov for a great help in the field.

References
Savage J. 2003. Revision of the genus Thricops Rondani (Diptera: Muscidae) // Insect Systematics and Evolution. Suppl.61. P.1 143. Sorokina V.S. 2012. Fauna of Muscidae (Diptera) of the Altai Mountains // Proceedings of the Russian Entomological Society, St. Petersburg. Vol.83. No.1. P.193222. Vikhrev N. 2011. Review of the Palaearctic members of the Lispe tentaculata species-group (Diptera, Muscidae): revised key, synonymy and notes on ecology // ZooKeys. Vol.84. P.5970. Vikhrev N., Sorokina V. 2009. Faunistic records of Thricops Rondani (Diptera, Muscidae) from Russia with description of two new species // Euroasian Entomological Jornal. Vol.8. No.3. P.341350 + III.