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Arthropoda Selecta 17(12): 101103

ї ARTHROPODA SELECTA, 2008

A new genus of the subfamily Ipainae from China (Arachnida: Aranei: Linyphiidae) Новый род подсемейства Ipainae из Китая (Arachnida: Aranei: Linyphiidae) Andrei V. Tanasevitch Андрей В. Танасевич
Centre for Forest Ecology and Productivity, Russian Academy of Sciences, Profsoyuznaya Str. 84/32, Moscow 117997 Russia. Email: atan@orc.ru. Центр по проблемам экологии и продуктивности лесов РАН, Профсоюзная ул. 84/32, Москва 117997 Россия.

KEY WORDS: Spiders, Linyphiidae, Ipainae, Ipaoides saaristoi, new genus, new species, China. КЛЮЧЕВЫЕ СЛОВА: Пауки, Linyphiidae, Ipainae, Ipaoides saaristoi, новый род, новый вид, Китай.

ABSTRACT. Ipaoides saaristoi gen.n., sp.n., is described from Yunnan, China. The genus belongs to the subfamily Ipainae Saaristo, 2007 and shows especially evident similarities to Uralophantes Esyunin, 1992. РЕЗЮМЕ. Описан новый род Ipaoides gen.n. с типовым видом Ipaoides saaristoi sp.n. из провинции Юньнань (Китай). Род входит в подсемейство Ipainae Saaristo, 2007 и наиболее близок к Uralophantes Esyunin, 1992.

Abbreviations
Abbreviations used in the text and figures: E embolus, MB median beam, MS membrosclerum, ME membraneous edge, R radix, RL radical lamella, PMP posterior median plate, WA wrinkled area, Tm I position of the trichobothrium on tibia I. Ipaoides gen.n.
ETYMOLOGY. The generic name refers to the subfamily name. Gender: masculine. DIAGNOSIS. Males are diagnosed by the long and coiled embolus unique among Ipainae, the presence of a large membrosclerum, as well as a forward directed dentiform extention in the distal part of the paracymbium. Females are diagnosed by the specific shape of the median beam of the epigyne. DESCRIPTION. Medium-sized ipaines, total length slightly exceeding 2 mm. Chaetotaxy unknown (see below). Metatarsus IV without trichobothrium. Palp (Figs 2 5): patella and tibia without special spines. Cymbium without posterodorsal outgrowth. Paracymbium L-shaped, with a distal dentiform extention directed forward. Membrosclerum (a heavily sclerotized median membrane transformed into a sclerite [see Tanasevitch, 1995]) large and infuscate. Embolic division highly complicated and characterized by several membraneous indistinct structures. Fickerts gland present, small, spherical. Radical lamella very long, slightly S-shaped. Embolus long, coiled, flat, with a wide, transparent, membraneous edge. Epigyne (Figs 69): large, protruded, a wrinkled area present. Median beam membraneous distally. Abdomen in both sexes with a dorsal pattern of two sets of paired spots, larger ones anterior. SPECIES INCLUDED. The type species only. DISTRIBUTION. The type locality only.
Type species Ipaoides saaristoi sp.n.

Introduction
The subfamily Ipainae Saaristo, 2007, was established for Ipa Saaristo, 2007, as the genotype [Saaristo, 2007]. Originally, Ipainae contained 29 species or subspecies from seven formerly micronetine genera. The present paper puts on record a new species from Yunnan, China, which belongs to a new genus of Ipainae.

Material and methods
This paper is based on material collected by Pekka T. Lehtinen (Turku, Finland) in Yunnan, China in 1987. The type specimens are housed in the Zoological Museum of Turku University, Turku, Finland (ZMTU). The terminology of the genitalic structures used here is after Saaristo [2007] and Tanasevitch [1995]. In the descriptions, the sequence of leg segments in measurement data is as follows: femur + patella + tibia + metatarsus + tarsus. All measurements are given in mm. Scale lines in figures = 0.1 mm.


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Figs 19. Ipaoides saaristoi sp.n.: 1 male body (dorsal view); 2 right palp; 3 paracymbium; 4 embolic division and membrosclerum (ventral view); 5 embolus; 6 female abdomen (ventral view); 79 epigyne (ventral, dorsal and lateral views, respectively). Abbreviations: E embolus, MB median beam, MS membrosclerum, ME membraneous edge, R radix, RL radical lamella, PMP posterior median plate, WA wrinkled area. Рис. 19. Ipaoides saaristoi sp.n.: 1 самец (вид сверху); 2 правая пальпа; 3 парацимбиум; 4 эмболюсный отдел и мембросклерум (вид снизу); 5 эмболюс; 6 абдомен самки (вид снизу); 79 эпигина (вид соответственно снизу, сверху и сбоку). Сокращения: E эмболюс, MB median beam, MS мембросклерум, ME membraneous edge, R радикс, RL radical lamella, PMP задняя медиальная пластинка, WA морщинистое поле.


A new genus of the subfamily Ipainae Saaristo
TAXANOMIC REMARKS. Based on the conformation of the secondary genital organs, the new genus shows special similarities to Uralophantes Esyunin, 1992, but differs distinctly by the more complicated structure of the embolic division, but mainly in the presence of a long, coiled, flat embolus. The epigyne of Ipaoides gen.n., in contrast to that of Uralophantes, has a posterior median plate.

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Ipaoides saaristoi sp.n. Figs 19.
MATERIAL. Holotype (ZMTU), China: Yunnan, Kunming Co, Songan, grassy hill slope, 13.X.1987, leg. P.T. Lehtinen. Paratype: 1 (ZMTU), same date and locality as in holotype.

above in the generic description. Abdomen 1.15 long, 0.70 wide, grey, dorsally with four white spots as in Fig. 1. Female. Known only from a crumpled abdomen. Abdomen 1.38 long, 0.85 wide, grey, with a readily distinguished dorsal pattern of large pale spots similar to that of the male. Epigyne (Figs 69): see above under the generic description. TAXONOMIC REMARKS. See above in the generic description. DISTRIBUTION. The type locality only. ACKNOWLEDGEMENTS. I am very grateful to Dr Pekka T. Lehtinen (Turku, Finland), who collected the material that served as the basis for the present paper, and to Dr Yuri M. Marusik (Magadan, Russia) who provided me with this material for study.

DIAGNOSIS. See above in the generic description. ETYMOLOGY. The species is named in memory of M.I. Saaristo, a well-known Finnish arachnologist, the author of fundamental publications devoted to linyphiid spiders. DESCRIPTION. Male. Total length 2.15. Carapace unmodified, 1.00 long, 0.75 wide, brown. Chelicerae 0.38 long. Legs yellow. The holotype specimen has only femora and one undamaged leg IV. Fe I, 1.13 long. Leg IV, 3.38 long (1.05+0.20+1.00+1.00+0.58). Chaetotaxy unknown. Metatarsus IV without trichobothrium. Palp (Figs 25): see

References
Saaristo M.I. 2007. A new subfamily of linyphiid spiders based on a new genus created for the keyserlingi-group of the genus Lepthyphantes (Aranei: Linyphiidae) // Arthropoda Selecta. Vol.16. No.1. P.3342. Tanasevitch A.V. 1995. Two new genera of the family Linyphiidae from Tuva, South Siberia, Russia (Arachnida Aranei Linyphiidae) // Ibid. Vol.4. No.3/4. P.6569.