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Arthropoda Selecta 19(2): 101-102

ї ARTHROPODA SELECTA, 2010

Inermocoelotes Ovtchinnikov, 1999, a valid genus (Aranei: Amaurobiidae: Coelotinae) Inermocoelotes Ovtchinnikov, 1999 валидный род (Aranei: Amaurobiidae: Coelotinae) Kirill G. Mikhailov К.Г. Михайлов
Zoological Museum, Moscow State University, Bolshaya Nikitskaya Str. 6, Moscow 125009 Russia. E-mail: mikhailov2000@gmail.com Зоологический музей МГУ, ул. Большая Никитская, 6, Москва 125009 Россия.

KEY WORDS: spiders, Coelotinae, new synonymy. КЛЮЧЕВЫЕ СЛОВА: пауки, Coelotinae, новая синонимия. ABSTRACT: A new synonymy is established: Eurocoelotes Wang, 2002 = Inermocoelotes Ovtchinnikov, 1999, syn.n. The rank of the latter taxon is elevated tothe genus level. Twelve new combinations are established. РЕЗЮМЕ: Установлена новая синонимия: Eurocoelotes Wang, 2002 = Inermocoelotes Ovtchinnikov, 1999, syn.n. Ранг последнего таксона поднят до родового. Установлено 12 новых комбинаций. The taxonomic position and rank of the group of genera related to Coelotes Blackwall, 1841, one is not clear. Before Lehtinen's [1967] revision, the genus and its relatives had usually been cited in Agelenidae sensu lato. Lehtinen definitely established a subfamily Coelotinae within Agelenidae. Most recent authors consider this subfamily within Amaurobiidae, but Nishikawa [2009] gave it a family rank, Coelotidae. The "intermediate" Amaurobiidae: Coelotinae concept is accepted herein. Currently, this group includes 583 species [Wang, 2010]. In his review of the ex-USSR Coelotinae taxonomy, the late S.V. Ovtchinnikov [1999] established three subgenera within the genus Coelotes: Coelotes s.str., Inermocoelotes subgen.n., and Brignoliolus subgen.n. These subgenera were overlooked in the later detailed revision of Coelotinae by Wang [2002], despite the fact that Ovtchinnikov's paper was cited in the revision. Brignoliolus Ovtchinnikov, 1999, definitely corresponds to Wang's charitonovi-group of Coelotes. One more conformity is discussed below. It is important to note that the Internet World Spider Catalog by Platnick [2010] does not include subgenera, even though nomenclatorial rules for subgenera are the same as those for genera [ICZN, 1999: rule 42.2]. The search for spider subgenera is difficult; only the respective original descriptions and several local catalogues can be recommended. By original designation, the subgenus Inermocoelotes Ovtchinnikov, 1999 included 6 species: C. inermis (L. Koch, 1855), as subgenerotype, C. drenskii Deltshev, 1990, C. karlinskii (Kulczyсski, 1906), C. kulczynskii Drensky, 1917, C. jurinitschi Drensky, 1915, and C. microlepidus De Blauwe, 1973. Four more species, C. titaniacus Brignoli, 1977, C. anoplus Kulczyсski, 1897, C. gasperinii Simon, 1891, and C. falciger Kulczyсski, 1897 are close to this subgenus. The genus Eurocoelotes Wang, 2002, originally included 11 species: E. inermis (L. Koch, 1855) as generotype, E. anoplus (Kulczyсski, 1897); E. brevispinus (Deltshev et Dimitrov, 1966); E. deltshevi (Dimitrov, 1993); E. drenskii (Deltshev, 1990); E. falciger (Kulczyсski, 1897); E. gasperinii (Simon, 1891); E. jurinitschi (Drensky, 1915); E. karlinskii (Kulczyсski, 1906); E. kulczynskii (Drensky, 1917); and E. microlepidus (Blauwe, 1973). Coelotes titaniacus Brignoli, 1977, was not included in Eurocoelotes. One other species, E. xinpingwangi Deltshev, 2009, was described recently. A comparison of the species listed in both Inermocoelotes and Eurocoelotes shows that all 6 species of Inermocoelotes are included in Eurocoelotes. Of the four species being close to Inermocoelotes, three also belong to Eurocoelotes. The type species is the same for both taxa. Therefore, the following synonymy is here established: Eurocoelotes Wang, 2002 = Inermocoelotes Ovtchinnikov, 1999, syn.n. The rank of Inermocoelotes is upgraded here to genus level. DIAGNOSIS. Differs from the closest genus Coelotes in having no patellar apophysis in the male and the presence of posteriorly situated epigynal teeth, large atrium and large copulatory ducts in the female. For a more detailed description, see Ovtchinnikov [1999: 74]. Wang [2002] provided no detailed description of the genus, but instead referred to the type species description only.


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K.G. Mikhailov
ACKNOWLEDGEMENTS. I am mostly grateful to Dr. P. Jдger for reviewing the ms. Drs Yu.M. Marusik and D.V. Logunov kindly commented on the ms helping to improve it. English of the final draft was kindly checked by Robin Leech.

COMPOSITION. The following species are included in Inermocoelotes Ovtchinnikov, 1999: Inermocoelotes inermis (L. Koch, 1855), comb.n. as the generotype. C- & S-Europe, Balkans. The easternmost records are Carpathians and Cis-Carpathia (both in Ukraine) and Moldavia. Reports of this species from Russian Plain (Lipetsk Area [Panteleeva, 1982], Voronezh Area [Panteleeva, Karavaev, 1992; Panteleeva, 2005]) are wrong. Inermocoelotes anoplus ( Kulczyсski , 1897), comb.n. Austria, Croatia, E-Europe. Inermocoelotes brevispinus (Deltshev et Dimitrov, 1966), comb.n. Bulgaria. Inermocoelotes deltshevi (Dimitrov, 1993), comb.n. Bulgaria. Inermocoelotes drenskii (Deltshev, 1990), comb.n. Bulgaria. Inermocoelotes falciger (Kulczyсski, 1897), comb.n. E-Europe, Bulgaria. Inermocoelotes gasperinii (Simon, 1891), comb.n. Croatia. Inermocoelotes jurinitschi (Drensky, 1915), comb.n. Bulgaria. Inermocoelotes karlinskii ( Kulczyсski , 1906), comb.n. SE-Europe. Inermocoelotes kulczynskii (Drensky, 1917), comb.n. Bulgaria. Inermocoelotes microlepidus (Blauwe, 1973), comb.n. Italy, Macedonia, Bulgaria. Inermocoelotes xinpingwangi (Deltschev, 2009), comb.n. Bulgaria. One other species, Coelotes titaniacus Brignoli, 1977, (Greece) probably belongs in this genus. DISTRIBUTION. Mainly the mountains and piedmonts of SE- and, in lesser extent, C-Europe.

References
ICZN. 1999. International Code of Zoological Nomenclature. Fourth ed. London: International Trust for Zoological Nomenclature. XXIX + 306 p. Lehtinen P. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha // Ann. Zool. Fenn. Vol.4. P.199-468. Nishikawa Y. 2009. A new genus and 44 new species of the family Coelotidae (Arachnidae, Araneae) from Japan // Ono H. (ed.). The spiders of Japan with keys to the families and genera and illustration of species. Kanagawa: Tokai Univ. Press. P.51-70. Ovtchinnikov S.V. 1999. [On the supraspeciefic systematics of the spider subfamily Coelotinae (Araneae, Amaurobiidae) in the former USSR fauna] // TETHYS Entomological Research. No.1. P.63-80 [in Russian, with English summary]. Panteleeva N.Yu. 1982. [To the study of spiders of the "Galichya Gora" Reserve] // Issledovaniya rastitel'nogo i zhivotnogo mira zapovednika "Galichya gora". Voronezh. P.89-92 [in Russian]. Panteleeva N.Yu. 2005. [Subphylum Chelicerata. Class Arachnida. Order Aranei] // Negrobov O.P. (ed.). Kadastr bespozvonochnykh Voronezhskoi oblasti. Voronezh. P.159-172 [in Russian]. Panteleeva N.Yu., Karavaev A.V. 1992. [Spiders of the Usmansky Bor Forest] // Sostoyanie i problemy ekosistem Usmanskogo bora. Voronezh. P.124-133 [in Russian]. Platnick N.I. 2010. The world spider catalog, version 10.5. American Museum of Natural History, online at http://research.amnh. org/iz/spiders/catalog/INTRO1.htm (accessed 23 April 2010) Wang Xin-Ping. 2002. A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae) // Bull. Amer. Mus. Nat. Hist. No.269. P.1-150. Wang Xin-Ping. 2010. Online Coelotinae, version 2.0. Online at http://www.amaurobiidae.com (accessed 24 April 2010)